Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
In this group we have two families of different age, the older, less
developed the Olenellidæ proper, and the younger the Paradoxidæ.
The former consists of the genera Olenellus, Holmia, Mesonacis
and Schmidtia, and we shall attend to them first. They have no
facial suture[6] and consequently a tripartite head shield like the
Archaic ones, no eyes, but there is that strongly developed and most
characteristic facial ridge. As seen in Olenellus Thompsoni[7] the
crescent like ridge starts as a direct outflow from the base of the
first segment of the glabella, and is in direct continuation with it
as an integral part. It is roundbacked, regularly curved and at its
starting point as broad as one of the posterior glabellar segments. It
is regularly semilunate, tapers posteriorly and ends near the occipital
segment. From the narrow second segment of the glabella, inside the
just described larger ridge, a smaller ridge emanates, broader and
flatter than the former, slightly curved, and ends between the third
and fourth glabellar segment. Already in Holmia Kjerulfi the ridge
is modified. There it is only one ridge, the anterior one, nearly as
doubled through a shallow groove running along its back. The second
one has dwindled away so as to be seen only as a narrow stripe near
the occipital segment and ending outside this in a point. The anterior
ridge, represented by HOLM as consisting of two nearly independent
parts, is indeed in one piece, though its dorsal groove sometimes is
deep, and it is with its total breadth joint to the first glabellar
segment. Along its outer edge, where it lies close to the cheek, a
narrow slit runs, and I suppose that it is to be considered as the
first indication of the forming of a facial suture, which however does
not reach longer than the ridge. As in Paradoxides it is so tight to
the cheek, that there has been no place for an eye. It is no accidental
break, its edges being too regular and unbroken in all specimens. This
ridge is by far much shorter than in Olenellus and terminates opposite
the third segment.
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