Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
In a little group that has retained larval or ancestral characters
during a great part of the palæozoic period, the genus Harpes stands
as a type. It has ranged from the oldest Lower Silurian, if we join
the related Harpides, to the middle Devonian. From near the top of
the glabella, though not so much forward as the facial ridge of the
blind trilobites (Olenus, Liostracus etc.) a straight ridge of much
varying length stands out on both sides and at its extremity two or
three globular stemmata with glossy surface lie encased. Probably this
ridge has the same origin as in the Olenidæ, the more so, as there are
indications of an extensive circulatory system. On plate IV fig. 18-19
the right hand ocelli of Harpes vittatus BARR. from Lochkow, Bohemia,
are represented. They are two, lying isolated near each other, quite
globular with circular outline, smooth and glossy as to exhibit a
shining surface. Their size is 0,4 mm in diameter. Being cut vertically
in the direction of the longitudinal axis of the head they resemble
elongated hemispheres, convex on the exterior surface, slightly concave
on the interior. The test of the head lies between them as a saddle and
covers them only partially and on the outer sides they lie with their
margins encased in the head shield. Seen in thin sections of the right
lens and magnified the whitish mass is traversed by vertical, blacker
streaks, standing somewhat radiating towards the sides and cancellate.
The other lens has a horizontal row of black dots. All this is
probably not of any structural value, only due to later changes.
The remarkable genus Harpides from the lowest Lower Silurian belongs
also to this group and has beside the fixed peduncle a peculiar
elongated ridge going from the eyes to the lateral margins of the
head[33], a ridge which is also present in some of the true Harpes.
[Footnote 33: Harpides breviceps cannot belong to this genus and is
rather related to Erinnys SALTER, as also MATTHEW holds it.]
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