Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
The hypostoma is square, with two short pointed wings, one on each side
of the slightly bent anterior margin and likewise two smaller ones
at the corners of the posterior margin. A groove follows on a short
distance the lateral margins and the posterior margin and disappears a
little below the anterior one. In the same direction, distally, though
a little more inside and unconnected with them there are two small
grooves, the bottom of which consists of a shell substance of different
colour and structure than the other parts. Having been a little ground
and seen in transmitted light it exhibits the shape of a club and a
homogenous yellow spot, tapering posteriorly and swelling out distally
(Pl. I fig. 4). It must be left an open question whether these maculæ
share in the nature of visual organs as the quite different maculæ of
the other trilobites, but it may be possible that it is so. It must,
however, be remarked that there are two types of hypostoma in the genus
Acidaspis as shown by the illustrations of BARRANDE. One has the small
grooves, possibly all sheltering the claviform maculæ, disposed as in
the now described A. crenata. This group embraces five species of the
Bohemian Silurian formation. The other group of three species again has
the hypostoma of the same quadratic or rectangular shape, but the two
short grooves, which may be expected to contain the maculæ, are placed
midways between the anterior and the posterior margins, nearly as the
maculæ bearing grooves of other trilobites. We have however not had
material for pursuing our researches in this genus, the other species
of the Swedish Acidaspidæ being unknown as to their hypostoma.
It may here be added an observation concerning the ornamentation of
the exterior surface of the hypostoma of Ac. crenata. It is covered
by a great number of diminutive circular or oblong wartlets occupying
the whole surface excepting the lower third of the central field just
above the posterior groove which is smooth. These wartlets seen through
transmitted light (Pl. I fig. 5) show in their interior something like
a peculiar black spiculum rising from a bifid rootlet and confined
within the wartlet and not extruding from it. In a longitudinal section
(Pl. I fig. 6) the spicula perforate the wartlets reaching through
their whole length. As in Calymmene these interior pseudo-spicula are
tubes, filled with iron-pyrites. It is probable that these tubes were
once bearing setæ and quite as in Apus formed a fur of bristles.
The structure of the cephalic eyes (Pl. 1 f. 1-2) is prismatic, but the
separate prisms are rather short and broad. Their lower or interior
end is convex. The separating lines between the single prisms are not
always distinct.
=Agnostus= DALM.
Pl. 1 fig. 7.
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