Speciation of the Wandering ShrewFindley, James S. (James Smith)
Science
Speciation of the Wandering Shrew
Findley, James S. (James Smith)
Shrews
populations which seem to represent intergrades between the two
subspecies. Isolated areas of Canadian Life-zone, even though
surrounded with Transition Life-zone, often harbor a population of
_obscurus_, whereas the streams in the nearby dry valleys harbor
populations of _vagrans_. Farther south in the Rocky Mountain chain,
_obscurus_ seemingly intergrades regularly with _vagrans_. This
intergradation is seen in populations from several localities in Utah.
There the lower elevations west of the Wasatch and Uinta mountains
are inhabited by _S. v. vagrans_, the higher elevations by _obscurus_
and where the ranges of the two abut intergrading populations occur.
In these series of intergrades there are specimens which, using size
as a subspecific criterion, would unhesitatingly be assigned, as
individuals, to _obscurus_, and others would be assigned to _vagrans_,
but these individuals represent extremes of a normally variable
population. At Cuddy Mountain, Idaho, the two subspecies seemingly
abut without intergradation; anyhow the available specimens from this
locality are referable to one or the other subspecies and none is
intermediate. The situation just described understandably has been the
source of much anguish to students who sought to identify shrews from
the Rocky Mountains. The reason for the relationship just described
has been discussed at length in a previous section.
In the Rocky Mountains of Wyoming and Colorado the subspecies _S. v.
obscurus_ ranges almost uninterruptedly over relatively large areas,
but southward in New Mexico and southwestward into Utah and Arizona,
suitable boreal habitat becomes insular in nature and obscurus there
is confined to the higher mountains. With one exception, once the
shrew populations become 'insular' in this region they become smaller
and show intergradation with _Sorex vagrans monticola_. The exception
is the population in the Sacramento Mountains of southeastern New
Mexico which is larger than _obscurus_ and has been rightly recognized
as a distinct subspecies, _neomexicanus_.
Almost without exception the range of typical _Sorex vagrans obscurus_
is sympatric with that of _Sorex cinereus_, usually the subspecies _S.
c. cinereus_. So close is this correspondence that the presence of _S.
cinereus_ comes near to being a useful aid in identifying _S. v.
obscurus_. In areas where individuals of _obscurus_ show
intergradation with _vagrans_, _Sorex cinereus_ is absent or rare. The
implication is that as the species _S. vagrans_ approaches the size of
the species S. cinereus, competition between the two increases with
resultant displacement of _cinereus_.
_Specimens examined._--Total number, 982.
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