Speciation of the Wandering ShrewFindley, James S. (James Smith)
Science
Speciation of the Wandering Shrew
Findley, James S. (James Smith)
Shrews
_Type._--Adult female, skin and skull; No. 23525/30943, U. S. Biol.
Surv. Coll.; obtained on August 26, 1890, by Vernon Bailey and B. H.
Dutcher from near Timber Creek, 8200 ft., Lemhi Mts., 10 mi. SSW
Junction (now Leadore), Lemhi Co., Idaho.
_Range._--Mountainous interior of western North America from central
Alaska east across Yukon and southwestern Northwest Territories to
northeastern Alberta, south in the mountains through north-central and
western Washington, Idaho, western Montana, Wyoming, Utah, and
Colorado, into northern New Mexico.
_Diagnosis._--Size medium to small for the species; average and
extreme measurements of 9 topotypes are: total length, 110.3
(105-117); tail, 46.4 (42-50); hind foot, 13.1 (12.5-13.5). Color
grayish or brownish gray in summer, light grayish in winter.
_Comparisons._--For comparisons with _S. v. setosus_, _S. v.
longicauda_, _S. v. alascensis_ and _S. v. shumaginensis_ see accounts
of those subspecies. Paler and slightly larger than S. v. soperi.
Larger than the subspecies from central Montana herein described as
new. Smaller than _S. v. neomexicanus_. Averaging larger in all
dimensions than _S. v. monticola_ with which _obscurus_ intergrades in
northern New Mexico and northern Arizona. Larger than _S. v. vagrans_
with more grayish rather than reddish fresh summer pelage and light
gray rather than dark grayish-black fresh winter pelage.
_Remarks._--Intergradation of _S. v. obscurus_ with _S. v. setosus_,
_S. v. longicauda_, _S. v. alascensis_, and the new subspecies from
Montana takes place in the usual way with specimens from intermediate
localities being intermediate in size and color. However the
relationship of _S. v. obscurus_ and _S. v. vagrans_ (as the latter
subspecies is defined in this study) is rather complicated. In
southern British Columbia where the two subspecies come together a
situation of remarkable complexity prevails. Series from some
localities seem to represent intergrades between _obscurus_ and
_vagrans_; from other localities some specimens seem to be referable
to one and some to the other subspecies; from other localities all
specimens seem referable to one subspecies. A similar situation is
seen in specimens from northeastern Washington, northern and central
Idaho, and extreme western Montana. The region mentioned is one of
extensive interfingering of life-zones. In southern British Columbia
the main axes of the rivers, valleys and mountain ranges are north and
south. Most of the valleys are in the Transition Life-zone; the
forests are rather dry and of pine with more or less isolated
hydrosere communities about streams and ponds. These hydrosere
situations are the habitat of _Sorex vagrans_. Shrews from these
situations are usually referable to _vagrans_. The high ridges and
mountain ranges are usually in the Canadian Life-zone or higher and
most of the shrews referable to _obscurus_ come from such places.
Marginal localities with regard to life-zone produce most of the
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