Speciation of the Wandering ShrewFindley, James S. (James Smith)
Science
Speciation of the Wandering Shrew
Findley, James S. (James Smith)
Shrews
Within the _Sorex vagrans_ complex, the only characters of taxonomic
significance that I have detected are in size and color. It is true
that cranial proportions, such as relative size of rostrum, may change
from population to population, but these proportions seem to me to be
dependent upon actual size of the individual shrew as I shall
elsewhere point out. Of the cranial measurements here employed,
palatal length and least interorbital breadth are the most significant
and useful. Color in the _S. vagrans_ group seems to be in Orange and
Cadmium Yellow, colors 15 and 17 of Ridgway (1912). No specimens
actually possess these pure colors, but most colors in these shrews
are seen to be derived from the two mentioned by admixture of black
and/or neutral gray. In color designations an increase in neutral gray
is indicated by an increased number of prime signs ('), whereas
increase in black is indicated by progressive characters of the Roman
alphabet (_i_, _k_, _m_). Thus, 17''_k_ is grayer than 17'_k_ and
17''_m_ is blacker than 17''_k_. In subspecific diagnoses in this
report, color and size, and sometimes relative size, are the
characters usually mentioned.
[Illustration: FIG. 3. Two measurements (in millimeters)
reflecting tooth-wear plotted against one another. First year
and second year individuals of _Sorex vagrans vagrans_, all
taken in August at Willapa Bay, Washington, are completely
separated. Open circles represent teeth of second year shrews;
solid circles represent teeth of first year shrews.]
PELAGE CHANGE
In general, winter pelage is darker than summer pelage in these
shrews. Winter pelage comes in first on the rump and spreads caudad
and ventrad. The growth line of incoming hair is easily detected on
the fur side of the skin. Throughout the winter the color of the
pelage changes, often becoming somewhat browner, although no actual
molt takes place. This was noted by Dalquest (1944) who assumed that
the color change resulted from molt although he was unable to detect
actual replacement of hairs. Summer pelage usually comes in first on
the back or head and moves posteriorly and laterally. Time of molt
depends on latitude and altitude. Summer pelage may appear fairly late
in the season and may account for the anomalous midsummer molt noted
by Dalquest. Fresh pelages of summer and winter are best seen in first
year animals and are less variable than are worn pelages and hence are
used as the basis of color descriptions.
GEOGRAPHIC DISTRIBUTION AND VARIATION
Pacific Coastal Section
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