Based on the idea of a direct creation of all [34] living beings, the
genera were then accepted as the created forms. They were therefore
regarded as the real existing types, and it was generally surmised that
species and varieties owed their origin to subsequent changes under the
influence of external conditions. Even Linnaeus agreed with this view in
his first treatises and in his "Philosophical Botany" he still kept to
the idea that all genera had been created at once with the beginning of
life.
Afterwards Linnaeus changed his opinion on this important point, and
adopted species as the units of the system. He declared them to be the
created forms, and by this decree, at once reduced the genera to the
rank of artificial groups. Linnaeus was well aware that this conception
was wholly arbitrary, and that even the species are not real indivisible
entities. But he simply forbade the study of lesser subdivisions. At his
time he was quite justified in doing so, because the first task of the
systematic botanists was the clearing up of the chaos of forms and the
bringing of them into connection with their real allies.
Linnaeus himself designated the subdivisions of the species as
varieties, but in doing so he followed two clearly distinct principles.
In some cases his species were real plants, and the varieties seemed to
be derived from them by [35] some simple changes. They were subordinated
to the parent-species. In other cases his species were groups of lesser
forms of equal value, and it was not possible to discern which was the
primary and which were the derivatives.
These two methods of subdivision seem in the main, and notwithstanding
their relatively imperfect application in many single examples, to
correspond with two really distinct cases. The derivative varieties are
distinguished from the parent-species by some single, but striking mark,
and often this attribute manifests itself as the loss of some apparent
quality. The loss of spines and of hairs and the loss of blue and red
flower-colors are the most notorious, but in rarer cases many single
peculiarities may disappear, thereby constituting a variety. This
relation of varieties to the parent-species is gradually increasing in
importance in the estimation of botanists, sharply contrasting with
those cases, in which such dependency is not to be met with.
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