The genus _Melitæa_, for example, can be characterized by the
possession of 7-9 fleshy tubercles bearing hairy spines; the genus
_Argynnis_ may be distinguished by always having six hairy unbranched
spines on each segment, and the genus _Cethosia_ by two similar spines
on each segment; the genus _Vanessa_ shows sometimes as many as
seven branched spines; and the genus _Limenitis_ never more than two
branched blunt spines on each segment, and so forth. If we go further
into details it will be seen that the most closely related imagines,
as might indeed have been expected, likewise possess the most nearly
allied larvæ, whilst very small differences between the imagines are
also generally represented by corresponding differences in the larvæ.
Thus, for instance, the genus _Vanessa_ of Fabricius has been divided
into several genera by later authors. Of these sub-genera, _Grapta_,
Doubl. (containing the European _C.-album_, the American _Fabricii_,
_Interrogationis_, _Faunus_, _Comma_, &c.), is distinguished by the
fact that the larvæ not only possess branched spines on all the
segments with the exception of the prothorax, but these spines are
also present on the head; in the genus _Vanessa_ (_sensû strictiori_),
Doubl., the head and prothorax are spineless (_e.g._ _V. Urticæ_); in
the tropical genus _Junonia_, Hübn., which was also formerly (Godart,
1819[186]) united with _Vanessa_, the larvæ bear branched spines on all
the segments, the head and prothorax included.
It is possible to go still further and to separate two species of
_Vanessa_ as two new genera, although they have hitherto been preserved
from this fate even by the systematists most given to “splitting.”
This decision is certainly justifiable, simply because these species
at present stand quite alone, and the practical necessity of forming a
distinct genus does not make itself felt, and this practical necessity
moreover frequently comes into conflict with scientific claims: science
erects a new genus based on the amount of morphological difference, it
being quite immaterial whether one or many species make up this genus;
such an excessive subdivision is, however, a hindrance to practical
requirements, as the cumbrous array of names thereby becomes still
further augmented.
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