It is probable that the greatest refrigeration of Norway, Sweden, Scotland, Wales, the Vosges, and the Alps coincided very nearly in time; but when the Scandinavian and Scotch mountains were encrusted with a general covering of ice, similar to that now enveloping Greenland, this last country may not have been in nearly so glacial a condition as now, just as we find that the old icy crust and great glaciers, which have left their mark on the mountains of Norway and Sweden, have now disappeared, precisely at a time when the accumulation of ice in Greenland is so excessive. In other words, we see that in the present state of the northern hemisphere, at the distance of about 1500 miles, two meridional zones enjoying very different conditions of temperature may co-exist, and we are, therefore, at liberty to imagine some former alternations of colder and milder climates on the opposite sides of the ocean throughout the Pleistocene era of a compensating kind, the cold on the one side balancing the milder temperature on the other. By assuming such a succession of events we can more easily explain why there has not been a greater extermination of species, both terrestrial and aquatic, in polar and temperate regions during the glacial epoch, and why so many species are common to pre-glacial and post-glacial times. The numerous plants which are common to the temperate zones north and south of the equator have been referred by Mr. Darwin and Dr. Hooker to migrations which took place along mountain chains running from north to south during some of the colder phases of the glacial epoch.* (* Darwin, "Origin of Species" chapter 11 page 365; Hooker, "Flora of Australia" Introduction page 18 1859.) Such an hypothesis enables us to dispense with the doctrine that the same species ever originated independently in two distinct and distant areas; and it becomes more feasible if we admit the doctrine of the co-existence of meridional belts of warmer and colder climate, instead of the simultaneous prevalence of extreme cold both in the eastern and western hemisphere. It also seems necessary, as colder currents of water always flow to lower latitudes, while warmer ones are running towards polar regions, that some such compensation should take place, and that an increase of cold in one region must to a certain extent be balanced by a mitigation of temperature elsewhere.