The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
[Illustration: Fig. 5.--A sketch of the so-called exites of _Neolenus
serratus_ (Rominger), to show the form and the character of the
spines. × 2.]
[Illustration: Fig. 6.--Endopodite of a cephalic appendage of
_Neolenus serratus_ (Rominger), showing the very broad coxopodite.
× 2.]
Before calling these structures new organs not previously seen on
trilobites, it is of course necessary to inquire if they can be
interpreted as representing any known structures. That they can not be
exopodites is obvious, since they are bordered by short stout spines
instead of setæ. The same stout spines that negate the above possible
explanation at once suggest that they are coxopodites (compare fig 6).
At first sight, the so-called exites seem too wide and too rounded to
be so interpreted, but if reference be had to the specimens rather
than the figures, it will be noted that the only well preserved
structure (No. 2) is longer than wide, has spines only on one side and
one end, and does not differ greatly from the coxopodite of specimen
No. 58589 (pl. 18, 1918). If structures 2, 1, and the segment ahead of
1 are really parts of one appendage, it can only be an endopodite, of
which No. 2 is the coxopodite, No. 1 the basipodite, and the next
segment the ischiopodite. If one looks carefully, there are no traces
of spines on either end of No. 1, but only on the margin. The extreme
width of No. 2 is against this interpretation as a coxopodite (see,
however, fig. 6), but it may be rolled out very flat, as this is an
unusually crushed specimen. No. 2 is 10 mm. long and 6 mm. wide at
the widest point. No. 1 is 5 mm. long and 3.5 mm. wide.
The crucial point in this determination is whether 2 and 1 are parts
of the same appendage. I believe they are, but others may differ.
Specimen No. 65513.
Illustrated: Walcott, Smithson. Misc. Coll., vol. 57, 1912, pl. 45,
fig. 3;--Ibid., vol. 67, 1918, pl. 16, figs. 1, 2.
This is nearly all of the right half of an entire specimen, but the
only appendages of any interest are those of the cephalon. Five
endopodites emerge from beneath that shield, but as all are displaced
it is not possible to say how many belong to the head. When held at
the proper angle to the light, the second and third from the front
show faintly the partial outlines of the coxopodites. The anterior
side and end of the best preserved one shows irregular stout spines of
unequal sizes, and the inner end is truncated obliquely (fig. 6).
These coxopodites are like those on the thorax of specimen No. 58589,
but shorter and wider. This of course suggests that the "exite" No. 2
of specimen No. 65515 may be a cephalic coxopodite. The endopodite of
this appendage, like the others on this cephalon, is shorter and
stouter than the thoracic or pygidial endopodites of the others
described.
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