The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
[Illustration: Fig. 7.--A restored section across the thorax of
_Neolenus serratus_, showing the probable form of attachment of the
appendages, their relation to the ventral membrane, and the jaw-like
endobases of the coxopodites.]
_Measurements:_ The cephalon is 24 mm. long and about 60 mm. wide. The
coxopodite of the third appendage is about 10 mm. long and 5.5 mm.
wide at the widest point. The corresponding endopodite is 19 mm. long
and projects 11 mm. beyond the margin, which is about 5 mm. further
than it would project were the appendage restored to its proper
position.
RESTORATION OF NEOLENUS.
(Text figs. 7, 8.)
This restoration is based upon the information obtained from the
studies which have been detailed in the preceding pages, and differs
materially from that presented by Doctor Walcott. The appendages are
not shown in their natural positions, but as if flattened nearly into
a horizontal plane. The metastoma is added without any evidence for
its former presence.
The striking features of the appendages are the broad unsegmented
exopodites which point forward all along the body, and the strong
endopodites, which show practically no regional modification. Although
the exopodites have a form which is especially adapted for use in
swimming, their position is such as to indicate that they were not so
used. The stout endopodites, on the other hand, probably performed the
double function of natatory and ambulatory legs.
[Illustration: Fig. 8.--_Neolenus serratus_ (Rominger). A restoration
of the ventral surface, with the endopodites omitted from one side, to
permit a better exposition of the exopodites. The position and number
of the appendages about the mouth are in considerable doubt. Restored
by Doctor Elvira Wood under the supervision of the writer. About
one-half larger than the average specimen.]
=Nathorstia transitans= Walcott.
Illustrated: Walcott, Smithson. Misc. Coll., vol. 57, 1912, pl. 28,
fig. 2.
The badly preserved specimen on which this genus and species was
based is undoubtedly a trilobite, but for some reason it does not
find a place in Walcott's recent article on "Appendages" (1918). The
preservation is different from that of the associated trilobites,
being merely a shadowy impression, indicating a very soft test. The
general outline of the body, the position of the eye, and even a
trace of spines about the pygidium (in the figure) are similar
to those of _Neolenus_, and I would venture the suggestion that
_Nathorstia transitans_ is a recently moulted _Neolenus serratus_,
still in the "soft-shelled" condition. Even if not a Neolenus, it is
probable, from the state of preservation, that it is an animal which
had recently cast its shell.
Walcott describes such fragments of appendages as remain, as follows:
Public-domain text, read in full here on John Shaqi.
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