The Appendages, Anatomy, and Relationships of TrilobitesRaymond, Percy E. (Percy Edward)
Science
The Appendages, Anatomy, and Relationships of Trilobites
Raymond, Percy E. (Percy Edward)
Trilobites
[Illustration: Fig. 23.--Transverse section of the thorax of _Calymene
senaria_, showing the large size of the mud-filled alimentary canal
(clear white). Traced from a photographic enlargement One appendifer
(also clear white) is shown. Specimen 153. × 3.3.]
_Ceraurus pleurexanthemus._
This species is taken up first, as it is the one shown in Walcott's
often-copied figure (1881, pl. 4, fig. 6). It is to be feared that too
many have looked at this figure without reading the accompanying
explanation, and have taken it for a copy of an actual specimen and
not a mere diagram, which it admittedly is. The evidence on which it
is based is comprised in eight transverse slices, one through the
glabella and seven through the thorax. Three of these have been
figured by Walcott: No. 27, 1881, pl. 3, fig. 7; No. 13, 1881, pl. 2,
fig. 3, 1918, pl. 26, fig. 14; No. 202, 1918, pl. 27, fig. 8. In all,
as can be seen by reference to the figures, the canal is partially
collapsed, and is much larger than is indicated in Walcott's
restoration. The other sections bear out the testimony of those
figured. One of these figured specimens (No. 27) and another figured
herewith (No. 118, see fig. 21) show an exceedingly interesting
structure which has previously escaped notice. The body cavity seems
to have had, in this region at least, a chitinous sheath on the dorsal
side. As shown especially in figure 21, this sheath impinges dorsally
and laterally against the axial lobe and thus furnishes a special
protection for the soft organs beneath, probably protecting them from
the strain of the dorsal muscles.
While there is no way in which the location of these sections in the
thorax can be positively determined, it is probable that they came
from the anterior end. In sections further back, supposed to be in the
posterior region of the mesenteron, no sheath is shown, but the canal
is nearly if not quite as large in relation to the size of the axial
lobe.
The single section through the glabella (specimen 97) is of course
important and fortunately well preserved (fig. 22). It shows the
dorsal sheath pressed against the inner surface of the axial lobe
along its middle portion, but diverging from it at the sides. The
section of the canal is oval, nearly twice as wide as high, but it is
obviously somewhat depressed. The original canal evidently filled
nearly the whole of the dorsal part of the glabella in this particular
region. Unfortunately, the connection with the mouth is not shown, and
the form of the hypostoma indicates that the section cut the glabella
diagonally, either in the anterior or posterior part, probably the
latter. In all these cases it should be remembered that the specimens
were found lying on their backs, and the canal has fallen in
(dorsally) since death.
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