Sladen (1912, _c_) also considers the possibility that the fluid which moistens the pollen might be secreted through the comb at the end of the tibia, through the tibio-tarsal joint, or from the surface of the auricle, but finds no evidence of glandular openings in these regions. A suggestion of a similar nature, apparently unknown to Sladen, was made by Wolff (1873), who describes "sweat-glands" which, he claims, are located within the hind tibia and the planta, and which pour a secretion upon the surface of the corbicula and upon the upper end of the planta through many minute openings located at the bases of hairs, particularly those which arise from the lateral margins of the corbicula. Wolff is convinced that the fluid thus secreted is the essential cohesive material by which the grains of pollen are bound together to form the solid mass which fills each fully loaded basket. He noticed that the mouthparts are used to collect pollen, and that some of it is moistened with "honey" or "nectar," but he does not consider that the fluid thus supplied is sufficient to explain adequately the facility with which the collecting bee brings together the scattered grains of pollen and packs them away securely in the baskets. Wolff's description of the basket-loading process itself is strikingly similar to that advocated later by Cheshire. The writer is not prepared to deny the possibility that the surface of the chitin of the hind legs of worker bees may be moistened by the secretion of glands which lie beneath it, but he is convinced that any fluid thus secreted bears little or no relation to the cohesion of the pollen grains within the baskets. Sections and dissected preparations of the hind legs of worker bees show certain large cells which lie within the cavity of the leg and which may function as secreting gland cells; but similar structures occur in even greater numbers within the hind legs of the drone and they are found within the hind legs of the queen. As has been noted, the extreme moisture of the plantar combs and of the tibio-tarsal articulation of the hind leg is readily understood when one recalls the manner in which moist pollen is compressed between the auricle and the tibial surface above it.