The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
MOTILE ORGANS.—Protoplasm often exhibits movements much more highly
specialised than the simple expansion or retraction of processes, or the
general change of form seen in Amoeba. If we imagine the activities of a
cell concentrated on particular parts, we may well suppose that they would
be at once more precise and more energetic than we see them in Amoeba or
the leucocyte. In some free-swimming cells, such as the individual cells
known as "Flagellata," the reproductive cells of the lower Plants, or the
male cells ("spermatozoa") of Plants as high as Ferns, and even of the
Highest Animals, there is an extension of the cell into one or more
elongated lash-like processes, termed "flagella," which, by beating the
water in a reciprocating or a spiral rhythm, cause the cell to travel
through it; or, if the cell be attached, they produce currents in the water
that bring food particles to the surface of the cell for ingestion. Such
flagella may, indeed, be seen in some cases to be modified pseudopodia. In
other cases part, or the whole, of the surface of the cell may be covered
with regularly arranged short filaments of similar activity (termed
"cilia," from their resemblance to a diminutive eyelash), which, however,
instead of whirling round, bend sharply {18}down to the surface and slowly
recover; the movement affects the cilia successively in a definite
direction in waves, and produces, like that of flagella, either locomotion
of the cell or currents in the medium. We can best realise their action by
recalling the waves of bending and recovery of the cornstalks in a
wind-swept field; if now the haulms of the corn executed these movements of
themselves, they would determine in the air above a breeze-like motion in
the direction of the waves (Fig. 5).[24] Such cilia are not infrequent on
those cells of even the Highest Animals that, like a mosaic, cover free
surfaces ("epithelium cells"). In ourselves such cells line, for instance,
the windpipe. One group of the Protozoa, the "Ciliata," are, as their name
implies, ciliated cells pure and simple.
[Illustration: FIG. 5.—Motion of a row of cilia, in profile. (From
Verworn.)]
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