The Cambridge natural history, Vol. 01 (of 10) — John Shaqi
The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
These, or some of them, may induce one of three different results, or a
combination thereof: (1) a single movement or an arrest of motion; (2) the
assumption of a definite position; (3) movement of a definite character or
direction.
{20}(i.) MECHANICAL STIMULI.—Any sudden touch with another body tends to
arrest all motion; and if the shock be protracted or severe, the retraction
of the pseudopodia follows. It is to this reaction that we must ascribe the
retracted condition of the pseudopodia of most Rhizopods when first placed
on the slide and covered for microscopic examination. Free-swimming
Protista may, after hitting any body, either remain in contact with it, or
else, after a pause, _reverse_ their movement, turn over and swim directly
away. This combination of movements is characteristic as a reaction of what
we may term "repellent" stimuli in general.[29] Another mechanical reaction
is that to continuous contact with a solid; and the surface film of water,
either at the free surface or round an air-bubble, may play the part of a
solid in exciting it; we term it "thigmotaxy" or "stereotaxy." When
positive it determines a movement on to the surface, or a gliding movement
along it, or merely the arrest of motion and prolongation of contact; when
_negative_, a contact is followed by the retreat of the being. Thus
_Paramecium_ (Fig. 55, p. 151) and many other Ciliates are led to aggregate
about solid particles or masses of organic _débris_ in the water, which
indeed serve to supply their food. On contact, the cell ceases to move its
cilia except those of the oral groove; as these lash backwards, they hold
the front end in close contact with the solid, at the same time provoking a
backward stream down the groove, which may bring in minute particles from
the mass.
(ii.) Most living beings are able to maintain their level in water by
floating or crawling against GRAVITY, and they react in virtue of the same
power against centrifugal force. This mode of irritability is termed
(negative) "geotaxy" or "barotaxy." We can estimate the power of resisting
such force by means of a whirling machine, since when the acceleration is
greater than the resistance stimulated thereby in the beings, they are
passively sent to the sides of the vessel. The Flagellates, _Euglena_ and
_Chlamydomonas_, begin to migrate towards the centre when exposed to a
centrifugal force about equal to ½ G (G = 32.2 feet or 982 cm. per second);
they remain at the centre until the centrifugal force is increased to 8 G;
above that they yield to the force, and are driven passively to the sides.
The reaction ceases or is reversed at high temperatures.
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