The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
{21}(iii.) RHEOTAXY.—This is the tendency to move against the stream in
flowing water. It is shown by most Protists, and can be conveniently
studied in the large amoeboid plasmodia of the Myxomycetes, which crawl
against the stream along wet strips of filter paper, down which water is
caused to flow. Most animals, even of the highest groups, tend to react in
the same way; the energetic swimming of Fishes up-stream being in marked
contrast with their sluggishness the other way; and every student of
pond-life knows how small Crustacea and Rotifers, no less than Ciliates,
swim away from the inrush of liquid into the dipping-tube, and so evade
capture. (See Vol. II. p. 216.)
(iv.) The movements of many Protozoa are affected greatly by LIGHT. These
movements have been distinguished into "photopathic," _i.e._ to or from the
_position_ of greatest luminosity; and "phototactic," along the _direct
path_ of the rays.[30] Those Protozoa that contain a portion of their
cytoplasm, known as a "plastid" or "chromatophore" (see pp. 36, 39),
coloured by a green or yellow pigment are usually "phototactic." They
mostly have at the anterior end a red pigment spot, which serves as an
organ of sight, and is known as an "eye-spot." In diffused light of low
intensity they do not exhibit this reaction, but in bright sunlight they
rise to the surface and form there a green or yellow scum.
Most of the colourless Protista are negatively phototactic or photopathic;
but those which are parasitic on the coloured ones are positively
phototactic, like their hosts.
Here, as in the case of other stimuli,[31] the absolute intensity of the
light is of importance; for as it increases from a low degree, different
organisms in turn cease to be stimulated, and {22}then are repelled instead
of being attracted. The most active part of the spectrum in determining
reactions of movement are the violet and blue rays of wave-length between
40 µ/10 and 49 µ/10, while the warmer and less refractive half of the
spectrum is inert save in so far as it determines changes in the
temperature of the medium.
(v.) The movements of many Protozoa are rendered sluggish by cold, and
active by a rise of TEMPERATURE up to what we may term the "optimum"; the
species becomes sluggish again as the temperature continues to rise to a
certain point when the movements are arrested, and the being is said to be
in a state of "heat-rigor." Most Protozoa, again, tend to move in an
unequally heated medium to the position nearest to their respective optimum
temperature. This is called "thermotaxy." The temperature to which Amoeba
is thermotactic is recorded as 35° C. (95° F.); that of _Paramecium_ is 28°
C. (82° F.).
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