The Cambridge natural history, Vol. 06 (of 10)Sharp, David
Science
The Cambridge natural history, Vol. 06 (of 10)
Sharp, David
Animals
_Sphecodes_ is a genus that has been the subject of prolonged difference of
opinion. The species are rather small shining bees, with a red, or red and
black, abdomen, almost without pollen-collecting apparatus, and with a
short but pointed ligula. These characters led to the belief that the
Insects are parasitic, or, as they are sometimes called, cuckoo-bees. But
evidence could not be obtained of the fact, and as they were seen to make
burrows it was decided that we have in _Sphecodes_ examples of industrial
bees extremely ill endowed for their work. Recent observations tend,
however, to prove that _Sphecodes_ are to a large extent parasitic at the
expense of bees of the genera _Halictus_ and _Andrena_. Breitenbach has
taken _S. rubicundus_ out of the brood-cells of _Halictus quadricinctus_;
and on one of the few {23}occasions on which this bee has been found in
Britain it was in circumstances that left little doubt as to its being a
parasite of _Andrena nigroaenea_. Marchal[21] has seen _S. subquadratus_
fight with _Halictus malachurus_, and kill it previous to taking possession
of its burrows; and similar observations have been made by Ferton. As the
older observations of Smith, Sichel, and Friese leave little doubt that
_Sphecodes_ are sometimes industrial bees, it is highly probable that we
have in this genus the interesting condition of bees that are sometimes
parasitic, at other times not; but so much obscurity still prevails as to
the habits of _Sphecodes_ that we should do well to delay accepting the
theories that have been already based on this strange state of matters.[22]
Friese states that in _Sphecodes_ the first traces of collecting apparatus
exist; and, accepting the condition of affairs as being that mentioned
above, it is by no means clear whether we have in _Sphecodes_ bees that are
abandoning the parasitic habit or commencing it; or, indeed, whether the
condition of uncertainty may not be a permanent one. It is difficult to
decide as to what forms are species in _Sphecodes_ owing to the great
variation. The Hymenopterist Forster considered that 600 specimens
submitted to him by Sichel represented no less than 140 species, though
Sichel was convinced that nearly the whole of them were one species, _S.
gibbus_. It has recently been found that the male sexual organs afford a
satisfactory criterion. The position of _Sphecodes_ in classification is
doubtful.
[Illustration: Fig. 11.—_Sphecodes gibbus_ ♀. Britain.]
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