Such was the theory of Lamarck, and it is sound and plausible in all
respects save one, namely, the unwarranted assumption that acquired
adaptations are inheritable, since these, to quote the words of the
Harvard zoölogist, G. H. Parker, “are as a matter of fact just the
class of changes in favor of the inheritance of which there is the
least evidence.” (“Biology and Social Problems,” 1914, p. 103.)
The next contribution to the philosophy of transformism was made by
Charles Darwin, when, in the year 1859, he published his celebrated
“Origin of Species.” In this work, the English naturalist bases the
evolution of organic species upon the assumed spontaneous tendency
of organisms to vary minutely from their normal type in every
possible direction. This spontaneous variability gives rise to slight
variations, some of which are advantageous, others disadvantageous to
the organism. The enormous fecundity of organisms multiplies them in
excess of the available food supply, and more, accordingly, are born
than can possibly survive. In the ensuing competition or struggle for
existence, individuals favorably modified survive and propagate their
kind, those unfavorably modified perish without progeny. This process
of elimination Darwin termed natural selection. Only individuals
favored by it were privileged to propagate their kind, and thus it
happened that these minute variations of a useful character were seized
upon and perpetuated “by the strong principle of _inheritance_.” In
this way, these slight but useful modifications would tend gradually
to accumulate from generation to generation in the direction favored
by “natural selection,” until, by the ensuing summation of innumerable
minor differences verging in the same direction, a major difference
would be produced. The end-result would be a progressive _divergence_
of posterity from the common ancestral type, whence they originally
sprang, ending in a multiplicity of new forms or species, all differing
to a greater or lesser extent from the primitive type. The contrary
hypothesis of a possible _convergence_ of two originally diverse types
towards eventual similarity Darwin rejected as an extremely improbable
explanation of the observed resemblance of organic forms, which, not
without reason, he thought it more credible to ascribe to their assumed
divergence from a common ancestral type.
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