Evolutionists in general, as we have said, regard our present plants
and animals as the modified progeny of earlier forms, understanding
by “modified” that which is the product of a trans-specific,
as distinguished from a varietal or intra-specific, change. To
substantiate the claim that changes of specific magnitude have actually
taken place, they appeal to two principal kinds of evidence, namely:
(a) empirical evidence based on such variations as are now observed
to occur among living organisms; (b) inferential evidence, which
aposterioristically deduces the common ancestry of allied organic
types from their resemblances and their sequence in geological time.
Hence, if we omit as negligible certain subsidiary arguments, the
whole evidence for organic evolution may be summed up under three
heads: (1) the genetic evidence grounded on the facts of variation;
(2) the zoölogical evidence based on homology, that is, on structural
resemblance together with all further resemblances (physiological and
embryological), which such similarity entails; (3) the palæontological
evidence which rests on the gradual approximation of fossil types to
modern types, when the former are ranged in a series corresponding to
the alleged chronological order of their occurrence in the geological
strata. It is the bearing of recent genetical research upon the first
of these three lines of evidence that we propose to examine in the
present chapter, an objective to which a brief and rather eclectic
historical survey of evolutionary thought appears to offer the easiest
avenue of approach.
While many bizarre speculations on the subject of transformism had
been hazarded in centuries prior to the nineteenth, the history
of this conception, as a scientific hypothesis, dates from the
publication of Lamarck’s “Philosophie Zoologique” in 1809. According
to Lamarck, organic species are changed as a result of the _indirect_
influence of the external conditions of life. A change in environment
forces a change of habit on the part of the animal. A change in the
animal’s habits results in adaptation, that is, in the development or
suppression of organs through use or disuse. The adaptation, therefore,
thus acquired was not directly imposed by the environment, but only
indirectly—that is, through the mediation of habit. Once acquired
by the individual animal, however, the adaptation was, so Lamarck
thought, taken up by the process of inheritance and perpetuated by
being transmitted to the animal’s offspring. The net result would be a
progressive differentiation of species due to this indirect influence
of a varying environment.
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