Thus organs of different animals are said to be homologous when they
are composed of like parts arranged in similar relation to one another.
Homologous organs correspond bone for bone and tissue for tissue, so
that each component of the one finds its respective counterpart in
the other. The organs in question may be functionally specialized
and externally differentiated for quite different purposes, but the
superficial diversity serves only to emphasize, by contrast, the
underlying identity of structure which persists intact beneath it.
Thus, for example, the wing of a pigeon, the flipper of a whale, the
foreleg of a cat, and the arm of a man are organs differing widely in
function as well as outward appearance, but they are called homologous,
none the less, because they all exhibit the same basic plan, being
composed of similar bones similarly disposed with respect to one
another.
Organs, on the other hand, are called analogous which, though
fundamentally unlike in structure, are, nevertheless, superficially
modified and specialized for one and the same function. The wing of
a bird and the wing of an insect furnish a trite instance of such
analogy. Functionally they subserve the same purpose, but structurally
they bear no relation to each other. In like manner, though both are
devoted to the same function, there exists between the leg of a man and
the leg of a spider a fundamental disparity in structure.
At times, specialization for the selfsame function involves the
emergence of a similar modification or uniform structural adaptation
from a substrate of basic dissimilarity. In these instances of parallel
modifications appearing on the surface of divergent types, we have
something more than mere functional resemblance. Structure is likewise
involved, albeit superficially, in the modification which brings
about this external uniformity. In such cases, analogy is spoken of
as _convergence_, a phenomenon of which the mole and the mole-cricket
constitute a typical example. The burrowing legs of the insect are,
so far as outward appearance goes, the exact replica on a smaller
scale of those of the mole, though, fundamentally, their structure is
quite unlike, the mole being built on the endoskeletal plan of the
vertebrates, whereas the mole-cricket is constructed on the exoskeletal
plan characteristic of the arthropods. Speaking of the first pair of
legs of the mole-cricket, Thomas Hunt Morgan says: “By their use the
mole-cricket makes a burrow near the surface of the ground, similar
to, but of course much smaller than, that made by the mole. In both
of these cases the adaptation is the more obvious, because, while the
leg of the mole is formed on the same general plan as that of other
vertebrates, and the leg of the mole-cricket has the same fundamental
structure as that of other insects, yet in both cases the details of
structure and the general proportions have been so altered that the
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