leg is fitted for entirely different purposes from those to which the
legs of other vertebrates and other insects are put.” (Quoted by Dwight
in “Thoughts of a Catholic Anatomist,” p. 235.) In the analogies of
convergence, therefore, we have the exact converse of the phenomenon so
often encountered in connection with homology. The latter exhibits a
contrast between basic identity and superficial diversity, the former a
contrast between superficial convergence and fundamental divergence.
Now the extreme importance of homology is manifest from the fact that
the taxonomists of zoölogy and botany have found it to be the most
satisfactory basis for a scientific classification of animals and
plants. In both of these sciences, organisms are arranged in groups
according as they possess in common certain points of resemblance
whereby they may be referred to this, or that, general type. The
resemblance is most complete between members of the same species, which
do not differ from one another by any major difference, though they may
exhibit certain minor differences justifying their subdivision into
varieties or races. These morphological considerations, however, must,
in the case of an organic species, be supplemented by the additional
physiological criteria of perfect sexual compatibility and normal
viability, as we have already had occasion to note in the previous
chapter. When organisms, though distinguished from one another by
some major difference, agree, notwithstanding, in the main elements
of structure, the several species to which they belong are grouped
under a common genus, and similarly genera are grouped into families.
A _relative_ major difference, such as a difference in the size of
the teeth, suffices for the segregation of a new species, while an
_absolute_ difference, such as a difference in the number of teeth or
the possession of an additional organ, suffices for the segregation of
a new genus. In practice, however, the classifications of systematists
are often very arbitrary, and we find the latter divided into two
factions, the “lumpers” who wish to reduce the number of systematic
groups and the “splitters” who have a passion for breaking up larger
groups into smaller ones on the basis of tenuous differences. Above
the families are the orders, and they, in turn, are assembled in still
larger groups called classes, until finally we reach the phyla or
branches, which are the supreme categories into which the plant and
animal kingdoms are divided. As we ascend the scale of classification,
the points of resemblance between the organisms classified are
constantly decreasing in number, while the points of difference
increase apace. Hence, whereas members of the same species have very
much in common, members of the same phylum have very little in common,
and members of different phyla show such structural disparity that
further correlation on the basis of similarities becomes impossible
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