The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
The Amphibia whose remains have been brought to light from the Coal
Measures have hitherto been regarded as pertaining to a single order, the
Stegocephalia, characterized by the completely roofed-over cranium and a
large parasphenoid. The writer (469) had previously assigned 5 suborders
to the group: the Branchiosauria, Microsauria, Aistopoda, Temnospondylia,
and Stereospondylia. All of these groups are represented in the Coal
Measures of North America. It has seemed inadvisable, in the light of our
present knowledge of the Amphibia, to retain these 5 groups as suborders,
and, in the revised scheme of classification which has been published
elsewhere (469), they are given the rank of orders all excepting the
Aistopoda, which are now regarded by the writer as specialized Microsauria.
The recent Caudata are possibly represented in the North American Coal
Measures by forms which may be assigned tentatively to the Proteida.
Such forms as _Cocytinus gyrinoides_, _Hyphasma lævis_, and _Erierpeton
branchialis_ possibly represent this group in the Pennsylvanian. This
relationship is based chiefly on the structure of the hyobranchial
apparatus and on the general structure of the species. The three
above-mentioned species are, however, very insufficiently known, and
the relationship can hardly be regarded as more than suggested by the
characters which are at hand.
The Salientia, or frogs, may possibly have their ancestral type in _Pelion
lyelli_, the first known species from the Linton, Ohio, Coal Measures.
Oddly enough, among the hundreds of specimens collected later from this
horizon, not a fragment can be identified with this species. The type
specimen is unique, and although incomplete its characters are suggestive.
The Branchiosauria are represented in North America by four species:
_Micrerpeton caudatum_, _Eumicrerpeton parvum_, _Mazonerpeton
longicaudatum_, and _M. costatum_. Three other genera which occur in North
America have been placed (642) in this group, but they do not belong
there, for reasons given below. The branchiosaurs were salamander-like
in appearance. They were naked, with the exception of small ovoid scales
on the back and the chevron-shaped armature of the ventral surface, the
latter being almost universally present among the Paleozoic Amphibia. They
were adapted for life in the water for at least the early part of their
existence, as is shown by the possession of gills on many of the late
Carboniferous and early Permian forms of Europe. The group is, without
doubt, ancestral to the modern Caudata. No branchiosaurians have been
described elsewhere from so low in the geological series as those here
given and they are the first and only evidence of the occurrence of the
group in the western hemisphere.
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