The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
The Microsauria are represented in the Coal Measures by numerous
forms which are usually characterized as lizard-like animals with a
well-developed ventral scutellation. Other characters, such as the
possession of lateral-line grooves on the cranium, the arrangement of the
cranial elements, and the condition of the ribs, will be discussed further
on. The pectoral arch is well developed and is composed of five dermal
bones plus the regular skeletal elements. The skeletal membrane bones are
sculptured after the manner of those of the cranium. The bodies of the
animals were, in a few cases, covered with scales; but most of them appear
to have been completely naked, even the ventral armature being absent in
some cases. The ventral scutellation was especially strong and highly
developed in some of the forms; _e.g._, in the genera _Saurerpeton_ and
_Sauropleura_. The vertebræ are uniformly of the hour-glass or notochordal
type. This is so generally the case that the characters of the vertebræ
and ribs are taken as the chief diagnostic characters of the major
groups. Various peculiarities are seen among the Microsauria, such as the
development of horns in various genera which are, apparently, related.
The order seems to have gone completely out of existence during the early
Permian, and if their descendants continued on as reptiles, as has been
suggested (469), we do not know the intermediate stages.
The Aistopoda are without doubt specialized microsaurs, and, in the
opinion of the writer, are not entitled to separate rank. Some of these
forms reached a high degree of specialization. One American species has
the skeleton reduced to a long, slender head and a slender series of
elongate vertebræ, all other parts of the skeleton, even the ventral
armature, being absent. The proportions attained by this species,
_Phlegethontia linearis_ Cope, recall those of the coach-whip snake,
_Zamenis flagellum_ Shaw, of the western plains. Some of the so-called
Aistopoda have been credited by Fritsch with the possession of peculiar
clasping organs, "Kammplatten." Newberry has written of the discovery of
similar structures in the Ohio Coal Measures (498), but the statement of
the actual association of these "Kammplatten" needs confirmation. Dr. R.
H. Traquair wrote to the author under date of April 28, 1909:
"I maintain that the association of a bundle of 'Kammplatten' with
a specimen of _Ophiderpeton_ in the Bohemian gas coal was entirely
_accidental_. Of such pitfalls the paleontologist has to beware or serious
mistakes may be the consequence, as has happened more than once. I must,
however, publish a short paper on the Kammplatten, for I think I know what
they are now."
Fritsch, however, has very clearly figured a nearly complete specimen
of _Ophiderpeton_ (251, Bd. IV) as possessing the Kammplatten in place
near the cloaca, where he suggests they may have served the function of
accessory copulatory organs or claspers.
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