The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
Below the coal-seam, nodules of various shapes and sizes occur, but they
seldom contain fossils and never good ones. Occasionally, as at Mazon
Creek, fragments of plants adhere to the outside of the incrusting shale.
A single nodule may have adhering to it fragments of 4 genera of plants.
The fossiliferous nodules all occur above the coal and are most prolific
and abundant immediately above the seam, within the first 10 inches. In
the same stratum of shale with the nodules are found abundant impressions
of plants in the shale, often perfect fronds being uncovered. (See, in
this connection, Twenhofel and Dunbar, 1914, "Nodules with Fishes from the
Coal Measures of Kansas," Amer. Journ. Sci., XXXVIII, pp. 157-163.)
G. F. Matthew (408-413) has described numerous genera and species of
footprints, presumably amphibian, from the Carboniferous of Canada. The
impressions indicate small creatures for the most part. Other imprints
have been described by Logan, Dawson, Lyell, Marsh, Mudge, and Lea. Since
the present work is intended largely for a morphological review, only
passing notice can be given to the _ichnites_. The literature on the
"Ichnites" has been brought together in Hay's "Bibliography and Catalogue
of Fossil Vertebrata of North America," pp. 538-553. References since the
publication of Hay's catalogue (317) will be found in the bibliography at
the end of this work. Footprints are of interest in that they are the only
evidence we have of the occurrence of land vertebrates in the Devonian and
Mississippian of North America.
CHAPTER IV.
THE MORPHOLOGY OF THE COAL MEASURES AMPHIBIA.
The anatomy of the Coal Measures Amphibia presents many primitive types
of structure. Their organization represents a stage passed through
in the ontogeny of higher vertebrates. The animals are similar in a
general way, yet so diverse are the modifications which they have
suffered under different environmental conditions, that close scrutiny
is needed to discern the exact relationship of the forms. Our knowledge
of this relationship is based on the structures preserved, which are
largely skeletal, since little is known of the soft anatomy (471) of the
air-breathing vertebrates of the Coal Measures. The pubis is ossified in
the Paleozoic Amphibia later than the ischium and ilium; the carpus and
tarsus are cartilaginous; the vertebræ consist of a pleurocentrum and two
neurocentra, thus paralleling conditions in modern mammalian embryos.
[Illustration: Fig. 6.--Generalized figure of dorsum of an early amphibian
skull to show position of elements and terminology adopted in this work.
The outline is based on that of _Eryops_, but is in no way intended to
indicate that form.
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