The Coal Measures Amphibia of North America — John Shaqi
The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
_a. com_, anterior commissure of lateral-line canals; _com_, commissural
communication between infra- and supra-orbital lateral-line canals; _fr_,
frontal; _inf_, interfrontal; _inn_, internasal; _info_, infraorbital
lateral-line canal; _it_, intertemporal; _jl_, jugal lateral-line canal;
_j_, jugal; _lar_, lacrimal; _mx_, maxilla; _n_, nasal; _oc_, occiput;
_occ_, occipital cross-commissure of the lateral-line system; _or_,
orbit; _par_, parietal; _pof_, postfrontal; _pmx_, premaxilla; _pf_,
prefrontal; _po_, postorbital; _pp_, postparietal; _g_, quadrate; _qj_,
quadratojugal; _spo_, supraorbital lateral-line canal; _sq_, squamosal;
_spt_, supratemporal; _t_, temporal lateral-line canal; _tab_, tabulare.]
(_a_) The skull of the Coal Measures Amphibia has (fig. 6) essentially
the same structure in the different groups. It is largely formed of
bones of intramembranous origin, representing the _face_ bones of the
mammalian skull. The skull in life was doubtless a _chondrocranium_ with
the membrane bones laid down upon the cartilaginous box containing the
sense-organs, as in the sturgeon (_Acipenser_), where the surface bones of
the face were probably originally scales, which later became consolidated
into large bony scutes. The membrane bones of the early Amphibia may have
been originally derived from scales, but at present nothing is known of
this origin; doubtless the elements had an intramembranous origin in the
ancestors of the group. Judging from Credner's studies on the series
of specimens of _Branchiosaurus amblystomus_ Credner (187), the skull
bones do not ossify completely until relatively late in the life of the
individual. The skull in the youngest individual figured by Credner
(_op. cit._, Taf. XVI, fig. 1) seems to be largely cartilaginous, with
beginnings of separation into the skeletal elements. The manner and
time of development and ossification of the skull seems to proceed much
as it does in modern amphibians. The condition found in the skull of
_Cryptobranchus allegheniensis_ or _Necturus maculosus_ will represent
pretty accurately the condition of most of the Coal Measures Amphibia. The
face bones in certain forms were sculptured and cut by lateral-line canals.
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