The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
A median suture divides the skull into two equal regions dorsally. On
the sides of this median suture lie pairs of elements which are common
to all higher vertebrates. These elements are: the premaxillæ, nasals,
frontals, parietals, and post-parietals. All of these elements vary
somewhat in shape and slightly in arrangement, but always occupy the same
relative positions. To the side of these elements lie the prefrontal, the
postfrontal, the supratemporal, the squamosal, and tabulare, and occupying
the margin of the skull are the maxilla, the jugal, the quadratojugal, and
possibly the quadrate in a few forms. The parietal foramen occurs usually
within the parietal bone, but its position is subject to slight variations
and it may occur on the suture between the frontal and the parietal, or
even far posterior near the postparietal. The nostrils often lie well
forward and are included by the premaxillæ, nasals, and prefrontals. The
orbit is usually well posterior, but it may occur far forward. It is
bounded by the prefrontal, the frontal, the postfrontal, the post-orbital,
and the jugal. Sometimes the lacrimal is present and has been clearly
identified on the anterior margin of the orbit in a few cases.
(_b_) _Sclerotic plates_ often occur within the orbits, and are not
confined to any particular group, though they are quite constant among
the Branchiosauria. They are usually delicate, thin, broad plates which
evidently overlap and operate as in modern animals. The number varies, as
many as 30 occurring within the orbit of one branchiosaur. Between the
margin of the orbit and the sclerotic plates there often occur, in the
Branchiosauria (186) particularly, small scale-like particles which were
doubtless embedded in the heavy skin above the orbit during life.
(_c_) The _palate_ of the skull is very incompletely known, being
indicated in a very few cases. These specimens, however, show that
the characters of the palate were quite similar, if not identical, in
essential respects with the palate among the European species of the same
or slightly later time.
A large cultriform parasphenoid occupies the posterior portion of the
palate, on either side of which in some species lies the posterior
palatine foramen. On the sides of the anterior prolongation of the
parasphenoid lie the vomers (186), membranous bones often bearing minute
tubercular teeth, apparently adapted for crushing. The vomers and the
maxillæ, with sometimes the palatine, surround the anterior palatine
foramen, which is almost always present; sometimes, however, quite small.
The transverse or ectopterygoid unites the pterygoid, a broad plate of
thin bone, with the maxilla and jugal.
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