The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
(_i_) The _alimentary canal_ (text-fig. 7) is beautifully preserved
as a cast in three specimens of the American branchiosaur species
_Eumicrerpeton parvum_ Moodie (471) from the Mazon Creek beds. The nodules
which contain these interesting little fossils measure less than 3 inches
in long diameter. The fossil salamanders, about 30 mm. in length, are
preserved on their backs and occur as nearly as is possible in the center
of the nodule.
If it were not for the fact that the œsophagus became loosened and dropped
from its place shortly after death, the alimentary canal would be in
place and would immediately recall a freshly dissected specimen of a
recent salamander. The anterior end of the œsophagus lies obliquely across
the chest region with its tip pointing slightly downward. The length
of the œsophagus proper, in one specimen, is only about 3 mm. As it is
preserved, the œsophagus lies in a semi-sigmoid curve with the convexity
anterior, and enters the cardiac portion of the stomach by a gradual
constriction. The stomach is clearly preserved as a distinct sac-like
organ with two lobes which correspond to the cardiac and pyloric limbs.
It measures about 7 mm. in length by 2 mm. in its greatest diameter.
The muscular constriction which divides the organ into pyloric and
cardiac divisions occurs at a distance of 4 mm. from the upper end. The
pylorus is designated by a rather pronounced constriction which may be
partly accidental, although it recalls the pylorus of modern frogs. From
this constriction, which lies on the left side of the fossil, as it is
preserved, the duodenal portion of the intestine makes a straight course
posteriorly to near the anal region, where it takes a sharp bend and
curves back to run parallel with itself for the distance of 4 mm. In its
upward course the intestine enlarges, and practically the same enlargement
continues throughout the remainder of the course to the anus. At a
distance of 1 mm. from the anal end, the rectum dilates probably 0.125
mm. to form the cloaca. After the intestine has continued its parallel
course for the 4 mm., as above stated, it turns abruptly to the right for
a distance of 2 mm. It then runs posteriorly for a short distance, then
bends back and under itself to again make a double sigmoid curve, when at
a distance of 6 mm. from the anus it assumes a straight course, which it
continues to the end.
[Illustration: Fig. 7.--Alimentary canal of Coal Measures salamander as
illustrated by the smaller specimen of _Eumicrerpeton parvum_ Moodie, from
the Mazon Creek shales. × 3. Original in Yale University Museum. _a_,
anus; _dd_, duodenum; _in_, intestine; _l_, impression of liver(?); _oes_,
œsophagus; _st_, stomach.]
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