The Coal Measures Amphibia of North America — John Shaqi
The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
The anus lies at a level which is approximately that of the lower end
of the femur, which is preserved as an impression on the left side of
the fossil, thus agreeing in its position with that found in modern
Caudata. Lying inside the curve of the stomach and partly inclosed by the
œsophagus is a smooth area which may possibly represent the impression
of some of the accessory digestive glands, such as the liver. Occurring
in this smooth area are numerous fine lines which possibly represent the
impressions of blood-vessels; but they are so imperfectly preserved that
one can not be sure.
Representatives of several genera of the modern Caudata have been
dissected in order to make a direct comparison of the fossil alimentary
canal with that of the recent forms. The alimentary tract of _Desmognathus
fuscus_ Raf. from the vicinity of Ithaca, New York, resembles in a
marked degree that of the fossil form. The nearest approach to the
condition there represented is found, however, in an immature branchiate
individual, some 47 mm. in length, of _Diemyctylits torosus_ Esch., from
a fresh-water pond on Orcas Island in Puget Sound. The presence of this
species on the island is very suggestive. It is of extreme interest, too,
that the condition represented in the alimentary tract of the fossil
branchiosaurs should resemble so closely that of an immature rather than a
mature form.
(_j_) The vertebral column is clearly and readily separable into cervical,
dorsal, sacral, and caudal regions. The neck is always short, with from
5 to 10 vertebræ, cervical ribs often present. The dorsal region is not
long, but varies from 20 to 30 in the constituent vertebræ. There is a
single sacral vertebra not always to be readily distinguished from those
of the dorsal and anterior caudal series. The tail may be very short
or extremely long, with neural and hæmal spines elongate and flattened
into an oar-like appendage. The distal caudals are in some species
cartilaginous, apparently always so in the Branchiosauria.
(_k_) The _atlas_ and _axis_ are unknown among the American specimens, but
we are able to infer from the structure of the other vertebræ what this
must have been; and our inferences are partly confirmed by the conditions
existing in the European forms (187). The atlas, apparently, consisted of
a pair of neurocentral plates which are partly ossified, partly embedded
in cartilage, judging from the edges of the plates which have been
preserved. The centrum seems not to have been present in the atlas, or if
present it was only very slightly developed and quite free from the neural
pieces and largely embedded in cartilage. A fairly accurate picture of the
condition of the _atlas_ and _axis_ may be seen on examining a cow, pig,
or chick embryo (378) in the early stages of vertebral development, which
has been cleared by the Schultze method (Amer. Journ. Anat., VII, No. 4,
1908).
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