The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
We have learned that the food of the crayfish is made up of very
diverse substances, both animal and vegetable; but, so far as they
are competent to nourish the animal permanently, these matters all
agree in containing a peculiar nitrogenous body, termed _protein_,
under one of its many forms, such as albumen, fibrin, and the like.
With this may be associated fatty matters, starchy and saccharine
bodies, and various earthy salts. And these, which are the essential
constituents of the food, may be, and usually are, largely mixed up
with other substances, such as wood, in the case of vegetable food,
or skeletal and fibrous parts, in the case of animal prey, which are
of little or no utility to the crayfish.
The first step in the process of feeding, therefore, is to reduce the
food to such a state, that the separation of its nutritive parts,
or those which can be turned to account, from its innutritious, or
useless, constituents, may be facilitated. And this preliminary
operation is the subdivision of the food into morsels of a convenient
size for introduction into that part of the machinery in which the
extraction of the useful products is performed.
The food may be seized by the pincers, or by the anterior chelate
ambulatory limbs; and, in the former case, it is usually, if not
always, transferred to the first, or second, or both of the anterior
pairs of ambulatory limbs. These grasp the food, and, tearing it into
pieces of the proper dimensions, thrust them between the external
maxillipedes, which are, at the same time, {50} worked rapidly to
and fro sideways, so as to bring their toothed edges to bear upon
the morsel. The other five pairs of jaws are no less active, and
they thus crush and divide the food brought to them, as it is passed
between their toothed edges to the opening of the mouth.
As the alimentary canal stretches from the mouth, at one end, to the
vent at the other, and, at each of these limits, is continuous with
the wall of the body, we may conceive the whole crayfish to be a
hollow cylinder, the cavity of which is everywhere closed, though it
is traversed by a tube, open at each end (fig. 6). The shut cavity
between the tube and the walls of the cylinder may be termed the
_perivisceral cavity_; and it is so much filled up by the various
organs, which are interposed between the alimentary canal and the
body wall, that all that is left of it is represented by a system
of irregular channels, which are filled with blood, and are termed
_blood sinuses_. The wall of the cylinder is the outer wall of the
body itself, to which the general name of _integument_ may be given;
and the outermost layer of this, again, is the _cuticle_, which
gives rise to the whole of the exoskeleton. This cuticle, as we have
seen, is extensively impregnated with lime salts; and, moreover, in
consequence of its containing _chitin_, it is often spoken of as the
_chitinous cuticula_.
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