(d) _The Velarium._ Charybdea, like most of the Cubomedusæ, possesses a
velum-like structure around the opening of the bell cavity (Fig. 3, _v_).
The velarium is a thin muscular diaphragm, resembling the true velum in
position and essential structures, but differing from the velum in its
origin, and in the possession of diverticula from the gastro-vascular
system, the velar canals. Of these there are in C. Xaymacana very
regularly sixteen, four in each quadrant. Their outline is seen in Fig.
3 to be forked with small irregular accessory processes. As for its
origin, the velarium of the Cubomedusæ is commonly accounted to have
arisen by fusion of marginal lobes, as in the case of the velarium of the
Discomedusæ. Pending decisive ontological evidence, the slight notches
in the four perradii seen in Fig. 3 may perhaps be taken as slight
indications of a primitive unfused condition, but the question will be
brought up again when the vascular lamellæ are discussed.
(e) _The Frenula._ Just as the stomach is attached to the walls of
the subumbrella in the four perradii by the suspensoria, so in the
lower part of the bell cavity the velarium is attached to the wall
of the subumbrella in the perradii by four structures similar to the
suspensoria, the frenula velarii. The frenula, like the suspensoria,
resemble the brackets of a shelf, with the difference that in the case of
the frenula the bracket is above the shelf, their purpose being evidently
to keep the velarium stiff against the outflow of water produced by
the pulsations of the bell. According to the greater need of strength
in this case, we find the frenula stouter, more buttress-like than the
suspensoria. The gelatinous ridge that gives them the necessary firmness
is thickened so as to be triangular in section, as shown in Fig. 16
(_frn_).
(f) _Musculature._ As is general in medusæ, the muscular system, so
far as known, is restricted to the subumbrella. It has a very simple
arrangement, consisting of a continuous sheet of circular (_i. e._
horizontal) striated fibres, which is interrupted only in the four
perradii by the radially directed muscle fibres of the suspensoria and
the frenula. In each quadrant, between the muscle of the suspensorium
above and that of the frenulum below, in an area just internal to the
sensory niche, there lies a space free from muscle. This interruption
of the muscle layer is shown in Fig. 39. Under the head of musculature
belonging to the subumbrella must be included also the radial, or
longitudinal muscles at the bases of the pedalia, which were mentioned
before (Fig. 16, _mt_). The mouth lobes and proboscis also are highly
contractile and muscular.
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