(g) _Nerve Ring._ It is in the possession of a clearly defined nerve ring
that the Cubomedusæ differ from all other Scyphomedusæ whose nervous
system has been carefully studied. The nerve ring shows very plainly
on the surface of the subumbrella as a well-defined clear streak. Its
course is zig-zag or festoon-like. In the interradii, at the basis of the
tentacles, it lies not far from the bell margin. In the perradii it rises
to the level of the sensory clubs. This very striking arrangement is
understood at once when it is remembered that the sensory clubs represent
the four perradial primary tentacles, and were originally situated on
the margin. When all the rest of the margin grew down and away from the
four sensory clubs, fusing below them to form the present intact edge of
the bell, the four portions of the nerve ring that lay in the perradii
were left at the level of the sensory clubs, and the originally straight
nerve ring was thus bent into a bow in each quadrant. The finer structure
of the nerve will be treated of in the special part to be devoted to the
nervous system.
c. _Internal Anatomy._
6. _Stomach._ The shape of the stomach is approximately that of a
biconvex lens, as seen in Fig. 4, which represents a Charybdea cut in
halves longitudinally in the perradius. The lumen of the proboscis (the
buccal stomach according to Haeckel’s terminology) communicates directly
by a funnel-shaped enlargement with the stomach proper, or central
stomach of Haeckel. The term basal stomach is carried over by Haeckel
from the Stauromedusæ, where it has considerable significance, to the
Cubomedusæ, and applied to the upper part of the central stomach. In the
stalkless Cubomedusæ, however, it has no significance so far as actual
structure goes, and our knowledge of the development of the Cubomedusæ is
as yet too simple for us to say that the upper part of the main stomach
represents what remains of the basal stomach of an earlier pedunculated
stage.
Public-domain text, read in full here on John Shaqi.
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