The same figure (No. 44) shows a characteristic difference between the
marginal lamella of Tripedalia and that of Charybdea. While in Charybdea,
as Claus points out, the marginal lamella keeps at one level, just a
little above the bell margin, all the way round (except where disturbed
by the special modifications of the tentacles and the sensory clubs),
and never descends into the velarium, in Tripedalia on the other hand
it describes a sinuous course, following the outlines of the marginal
pockets, as is indicated in the figure by the light parallel line _vlm_.
The course as it would be seen in a surface view is obscured just at
each side of the interradius by the overhanging of the bases of the two
lateral pedalia. This is why the lamella is not indicated at these points
in the diagram. The course is seen to lie almost wholly on the velarium,
that is, in the figure below the line which represents the bell margin
proper, the line at which the angle comes when the velarium is in its
normal position, horizontal to the vertical side of the bell.
In this sinuous course of the marginal lamella we have another point of
resemblance between Tripedalia and the Chirodropidæ. H. V. Wilson worked
it out in his sections of Chiropsalmus, and the reconstruction which I
have given in the figure under discussion is in all essentials similar
to his for Chiropsalmus. The differences lie only in the fact that
Chiropsalmus has more velar canals, and that the chief marginal pocket
in each quadrant is not forked peripherally, as is that of Tripedalia
(_mp_), but presents its distal margin parallel to the edge of the
velarium. The two smaller marginal pockets in the perradii (_mp´_) are on
identically the same plan in both.
Tripedalia, having three tentacles joining the umbrella in each
interradius, shows a disturbance of the course of the marginal lamella in
these regions by just so much the more complicated than in Charybdea. The
plan, however, is exactly the same. The lamella is pushed upwards from
the margin by each of the bases of the three pedalia just as is done by
the base of the single pedalium of Charybdea. Fig. 29 shows the lamella
in the same relation to the canal of the central tentacle (_ct_) that
it has in the similar sections of Charybdea (Figs. 16 and 43); and in
addition the first appearances (as the series is traced downwards) of
the arches of the lamella over the two lateral tentacles (_ct´_), which
are inserted a little lower down than the middle one of the group. As
concerns these lateral tentacles, the relations of the vascular lamella
at this level are the same as that in the level of Fig. 40 for Charybdea.
Public-domain text, read in full here on John Shaqi.
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