It has been stated more than once already that the vascular lamella of
the sensory niche is a part of the lamella that runs round the margin,
and so far the only evidence given has been the strip of endodermal
fusion running from the marginal lamella to that of the niche. This
strip, however, as has been described, does not come to the surface and
consequently seems at first sight to be a different structure from the
lamella of the margin. That it is not, however, I found very prettily
shown in a series of sections of one of my youngest Tripedalia. In
this the lamella of the niche as it was traced in successive sections
downwards, was found not to form a closed ring at the bottom of the
niche, but each side was continued directly and separately downwards to
the margin, where it passed into the corresponding part of the marginal
lamella. A reconstruction of the condition, similar to that of Fig. 44,
is given in Fig. 45, and I think explains itself at a glance. Evidently
the vascular lamellæ that connect the lamella of the sensory niche with
that of the margin at first come to the surface, like the rest of the
marginal system, but as the animal grows older come to lie within the
gelatine. In this way the condition found in cross-sections just through
the margin of my very small Tripedalia, and represented in Fig. 46,
becomes that of the adult seen in the corresponding portion of Fig. 29.
It is as complete a demonstration as could be required that the lamella
of the sensory niche is at first only a loop of the marginal lamella, a
conclusion that had been already reached by H. V. Wilson on theoretical
considerations, based upon the facts of the adult structure as he found
them in Chiropsalmus.
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