The way in which the nerve reaches the sensory clubs is interesting.
Under the topic of the vascular lamellæ it was explained that the sensory
clubs and the bottom of the sensory niche from which they spring are
parts of the subumbrella. Fig. 37 reminds at a glance better than any
other one drawing how the bottom or inner wall of the niche is completely
cut off from the exumbrella by vascular lamellæ above and below the
stalk of the club. From this figure, now, it will readily be understood
that the nerve in order to pass to the base of the stalk has simply
to traverse the gelatine of the subumbrella. This fact, which seems
surprising enough at first sight in view of the position of the clubs
on the external surface of the umbrella, was correctly pointed out and
explained by Claus, but one or two figures will serve perhaps to give a
clearer idea of it.
Fig. 49 is a diagram of the nervous structures in the region of the
sensory niche, as they would be seen on the surface of the subumbrella
turned toward the bell cavity. The outline of the sensory niche as it is
seen through the tissue of the animal is represented by the line _osn_.
The sensory club (_scl_), and its stalk with a conical basal portion are
given by the lightly dotted outline and are also imagined as seen through
the animal. The nerve (_n_), being on the surface of the subumbrella, is
shown as a heavy line describing an arch over the outline of the niche.
In the middle point of the arch is a slight thickening of the nervous
tissue (_rg_) which shows in section a large increase in the number of
ganglion cells, and is the radial ganglion of Claus. The same is seen,
exaggerated in size, in Fig. 12. From it there extends upward a slender
strand of nervous tissue (_rn_), the radial nerve of Claus. In Charybdea
this can be traced but a very short distance. In Tripedalia it is much
more distinct and traceable for a longer distance, and I might say in
passing that this and the sensory organs in the proboscis are the only
differences I have noted between the nervous systems of Tripedalia and
Charybdea.
Nerve ring, radial ganglion and radial nerve all lie on the bell cavity
surface of the subumbrella. The way, now, in which the nerve ring reaches
the base of the stalk is simply by sending two roots through the gelatine
of the subumbrella to the conical base of the stalk. These roots are seen
in the diagram at _rns_. After passing through the gelatine the roots
come together on the inner side of the base--that is, the side turned
toward the bell cavity--and then pass downwards (_nst_) on the inner side
of the stalk of the club to the mass of nervous tissue at its end.
Public-domain text, read in full here on John Shaqi.
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