This passage of nervous tissue through the gelatine in order to reach the
sensory club is a little hard to grasp at the first, and I have tried
to render it more intelligible by a couple of drawings of sections.
Fig. 50 is a transverse section through the upper part of the region of
the sensory niche, not quite horizontal (_i. e._ parallel with the bell
margin), but slanting so as to lie on the plane of the reference arrow
_x-y_ in Fig. 49. The plane passes just through the top of the niche, and
in two areas has cut through the roof with its epithelium of ectoderm
(_ece_, _ecs_) so that the space of the sensory niche (_sn_) appears. The
vascular lamella of the sensory niche (_vls_) is shown, as in Figs. 13
and 14, running on each side from the endoderm that lines the canal of
the sensory club (_enc_) to the endoderm of the adjacent stomach pocket
(_enp_). By it the gelatine of the exumbrella is separated from that of
the subumbrella, and one sees that it is only through the latter that
the nerve has to pass in order to reach the base of the sensory club. It
is also seen that one part of the roof of the niche which is cut through
lies outside of the ring of lamella and is therefore lined with ectoderm
of the exumbrella (_ece_) while the other lies within the ring and is
lined with ectoderm of the subumbrella (_ecs_). Owing to the slanting
direction of the cut only the root on one side is cut through. The other
is indicated, however, on the right side of the drawing. In this method
of passage of nerve fibres, together with the accompanying ganglion
cells, directly through the gelatine to the stalk of the sensory club my
work is only confirmation and explanation of Claus.
Fig. 51 is a vertical section through the base of the stalk in the plane
of the reference arrow _w-z_ in Fig. 49, and therefore passing through
one of the roots of the nerve of the stalk. Here again the region is seen
to be cut off from the exumbrella by the vascular lamella of the sensory
niche (_vls_), and the nerve is seen passing through the gelatine of
the subumbrella from the surface of the bell cavity (_sc_) to the base
of the stalk hanging in the sensory niche (_sn_). One of the ganglion
cells (_gc_) that accompany the nerve is seen to have two nuclei, a not
infrequent occurrence which has been pointed out by others.
Public-domain text, read in full here on John Shaqi.
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