We saw that conduct is distinguished from the totality of actions by
excluding purposeless actions; but during evolution this distinction
arises by degrees. In the very lowest creatures most of the movements
from moment to moment made, have not more recognizable aims than have
the struggles of an epileptic. An infusorium swims randomly about,
determined in its course not by a perceived object to be pursued or
escaped, but, apparently, by varying stimuli in its medium; and its
acts, unadjusted in any appreciable way to ends, lead it now into
contact with some nutritive substance which it absorbs, and now
into the neighborhood of some creature by which it is swallowed and
digested. Lacking those developed senses and motor powers which higher
animals possess, ninety-nine in the hundred of these minute animals,
severally living for but a few hours, disappear either by innutrition
or by destruction. The conduct is constituted of actions so little
adjusted to ends, that life continues only as long as the accidents of
the environment are favorable. But when, among aquatic creatures, we
observe one which, though still low in type, is much higher than the
infusorium--say a rotifer--we see how, along with larger size, more
developed structures, and greater power of combining functions, there
goes an advance in conduct. We see how by its whirling cilia it sucks
in as food these small animals moving around; how by its prehensile
tail it fixes itself to some fit object; how by withdrawing its outer
organs and contracting its body, it preserves itself from this or that
injury from time to time threatened; and how thus, by better adjusting
its own actions, it becomes less dependent on the actions going on
around, and so preserves itself for a longer period.
A superior sub-kingdom, as the Mollusca, still better exemplifies this
contrast. When we compare a low mollusc, such as a floating ascidian,
with a high mollusc, such as a cephalopod, we are again shown that
greater organic evolution is accompanied by more evolved conduct.
At the mercy of every marine creature large enough to swallow it,
and drifted about by currents which may chance to keep it at sea, or
may chance to leave it fatally stranded, the ascidian displays but
little adjustment of acts to ends in comparison with the cephalopod;
which, now crawling over the beach, now exploring the rocky crevices,
now swimming through the open water, now darting after a fish, now
hiding itself from some larger animal in a cloud of ink, and using
its suckered arms at one time for anchoring itself and at another
for holding fast its prey; selects and combines and proportions its
movements from minute to minute, so as to evade dangers which threaten,
while utilizing chances of food which offer: so showing us varied
activities which, in achieving special ends, achieve the general end of
securing continuance of the activities.
Public-domain text, read in full here on John Shaqi.
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