But though we cannot directly follow the uninterrupted process of
natural selection which is going on under natural conditions, there is
another kind of proof for this hypothesis, besides that which consists
in logically deducing a process from correct premisses; I should like
to call this the practical proof. If a hypothesis can be made to
explain a great number of otherwise unintelligible facts, it thereby
gains a high degree of probability, and this is increased when there
are no facts to be found which are in contradiction to it.
Both of these criteria are fulfilled by the selection-hypothesis, and
indeed the phenomena which may be explained by it, and are intelligible
in no other way, present themselves to us in such enormous numbers,
that there can be no doubt whatever as to the correctness of the
principle; all that can be still disputed is, how far it reaches.
Let us now turn our attention to this practical way of proving the
theory by the facts which it serves to interpret, beginning with a
consideration of the external appearance of organisms, their colour and
form.
_The Colour and Form of Organisms._
Erasmus Darwin had in many cases already rightly recognized the
biological significance of the colouring of an animal species, and
we may be sure that many of the numerous good observers of earlier
times had similar ideas. I can even state definitely that Rösel von
Rosenhof, the famous miniature-painter and naturalist of Nürnberg in
the middle of the eighteenth century, recognized clearly, and gave
beautiful descriptions of what we now call colour-adaptation. It is
true that he gave them only as isolated instances, and was far from
recognizing the phenomenon of colour-adaptation in general, or even
from inquiring into its causes. From the time of Linné, the endeavour
to establish new species overshadowed all the finer observation of
life-habits and inter-relations, and, later on, after Blumenbach,
Kielmeyer, Cuvier, and others, the eager investigation of the internal
structure of animals also tended to divert attention from these
œcological relations. In systematic zoology, colour ranked only as a
diagnostic character of subordinate value, because it is often not very
stable, and indeed is sometimes very variable; it was therefore found
preferable to keep to such relatively stable differences as are to be
found in the form, size, and number of parts.
Charles Darwin was the first to redirect attention to the fact that the
colouring of animals is anything but an unimportant matter; that, on
the contrary, in many cases it is of use to the animal, e.g. in making
it inconspicuous; a green insect is not readily seen on green leaves,
nor a grey-brown one on the bark of a tree.
Public-domain text, read in full here on John Shaqi.
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