Now, if in the course of the phylogeny numerous new segments were
added to the body of the Crustacean, the ontogeny would be materially
prolonged, and condensation would become necessary in the interests of
species-preservation. To bring this condensation about, whole series of
segments which were added successively in the phylogeny succeeded each
other with gradually increasing rapidity in the ontogeny, until finally
they appeared _simultaneously_: the determinants of the segments _n_,
_n_ + 1, _n_ + 2, ... _n_ + _x_ varied in regard to their liberating
stimuli, and were roused to activity no longer successively, but
simultaneously, in the cell complexes controlled by them. We have thus
recapitulation, but with abridgement and compression, of the phyletic
stages in the ontogeny. Thus in the nauplius of _Leptodora_ we see the
rudiments of five of the pairs of legs of the subsequent thorax (Fig.
111, _IV-VIII_), and in the Zoæa larva the rudiments of six thoracic
legs may be seen behind the already developed swimming-leg (Fig. 114,
_VI-XIII_).
But in the course of the phylogeny a segment may also become
superfluous, and we know that it then degenerates and is ultimately
eliminated altogether. Thus in a parasitic Isopod, which lives
within other Crustaceans, a segment of the thorax is wanting in the
relatively well-developed larva, and in the Caprellidæ among the
Amphipod Crustaceans the whole abdomen of from six to seven segments
has degenerated to a narrow, rudimentary structure. In such cases
the gradual degeneration of the relative determinants has preceded
step for step the degeneration of the part itself, and when this is
complete the ontogeny shows nothing of what was previously present,
and so we may speak of a 'falsification' of the phylogeny. But that
the complete disappearance of the determinants only comes about with
extreme slowness, so that whole geological periods are sometimes not
enough for its accomplishment, we have already learnt from our study
of rudimentary organs, instances of which can be demonstrated in every
higher animal, bearing witness to the presence of the relevant organs
or structures in the ancestors of the species.
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