The Fern Bulletin, October 1903: A Quarterly Devoted to FernsVarious
Science
The Fern Bulletin, October 1903: A Quarterly Devoted to Ferns
Various
Ferns
Type European. British Columbia, near Wharnock Station, _A. J. Hill_;
Vancouver, _Macoun_ (as _ramosissimum_); Blacktail Deer Creek,
Yellowstone Park, _Knowlton_. The latter is quite peculiar in appearance
and approaches _robustum_. None of the specimens exactly agree, but will
come here better than elsewhere. _The Ames Botanic Laboratory, North
Easton, Mass._
THE SPECIES-CONCEPTION AMONG THE TERNATE BOTRYCHIUMS.
By Willard N. Clute.
Living as I do in the midst of a region rich in specimens of the ternate
_Botrychiums_, I have taken more than ordinary interest in the
discussion of the relative rank to which the various forms should be
assigned. After considerable study of the subject which has consisted of
a careful balancing of the degree of differentiation in each form, as
well as an examination of much material both in the herbarium and in the
field, I have come to certain conclusions which I purpose to set down
here.
Before the separate forms are discussed it may be well to say a few
words on the variations of _Botrychium ternatum_ in general. It is a
noticeable fact that all the so-called new species of this section of
the genus, have been based primarily upon the cutting of the sterile
part of the frond. This is all the more remarkable since there are
probably no other genera in which species are founded on the minor
outlines of a mere leaf. One has but to turn to nature in any clime to
see that leaves are not invariably of the same shape. Note the wide
variation in the moonseed, the hollyhock, the sassafras, and some of the
buttercups among flowering plants, and if it be contended that the cases
are not parallel, take as further illustration the blood-root, which,
like the _Botrychium_, produces but one leaf a year, and note the
cutting of its single leaf. If all these forms of _Botrychium_ are
species, why have not the forms of the bloodroot been segregated?
Moreover, if we are to recognize these forms of _Botrychium_ as species,
why should we not also recognize as such the three hundred forms of
_Athyrium filix-foemina_, or the hundred or more forms of
_Scolopendrium_? It is unavailing to say that these latter are mere
gardeners’ varieties, for we have it on the authority of Mr. Druery, who
is familiar with them all, that a large number come true from spores.
Experiments with flowering plants have shown that the thickness of
leaves and the amount of cutting of their edges, may be altered by
different degrees of moisture, sunshine, etc., to which they are
exposed, and we may infer as much for the ferns. This being so, it is
not difficult to account for the slight variations in cutting exhibited
in plants from widely separated points in the United States.
Public-domain text, read in full here on John Shaqi.
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