Base of wing-blade thin or slightly
thickened.
Cones dehiscent at maturity.
Pits of ray-cells large X. Lariciones
Pits of ray-cells small XI. Australes
Cones serotinous, pits of ray-cells small XII. Insignes
Base of wing-blade very thick XIII. Macrocarpae
HAPLOXYLON
Bases of the bracts subtending leaf-fascicles not decurrent. Staminate
flowers not sufficiently developed in the bud to be apparent.
Spring-shoots uninodal. Fibro-vascular bundle of the leaf single. Cone
symmetrical, of relatively fewer larger scales, its tissues softer.
Bark-formation late, the trunks of young trees smooth. Wood soft and
with little resin, of uniform color and with relatively obscure
definition of the annual rings. Tracheids of the medullary rays with
smooth walls.
All the species of this section, except P. Nelsonii, have deciduous
fascicle-sheaths. There are but two species of Diploxylon with
deciduous sheaths, P. leiophylla and P. Lumholtzii, both of them
easily recognized. The deciduous sheath, therefore, is an obvious and
useful means for recognizing the Soft Pines. On the characters of the
fruit and the wood Haploxylon can be divided into two subsections.
a. Cembra Umbo of the cone-scale terminal.
b. Paracembra Umbo of the cone-scale dorsal.
=Cembra=
Umbo of the cone-scale terminal. Scales of the conelet unarmed. Leaves
in fascicles of 5, the sheath deciduous, the two dermal tissues
distinct, the hypoderm-cells uniform. Pits of the cells of the wood-rays
large.
Seeds wingless.
Cones indehiscent I. Cembrae.
Cones dehiscent II. Flexiles.
Seeds with an adnate wing III. Strobi.
=I. CEMBRAE=
Seeds wingless. Cones indehiscent, deciduous at maturity.
In this group of species there is no segregation of sclerenchyma into
an effective tissue. The cones are inert under hygrometric changes and
may always be recognized in herbaria by their persistent occlusion and
soft tissues. The seeds are released only by the disintegration of the
fallen cone. There is, however, a vicarious dissemination by predatory
crows (genus Nucifraga) and rodents.
Leaves serrulate, their stomata ventral only.
Cones relatively larger, the apophyses protuberant 1. koraiensis.
Cones relatively smaller, the apophyses appressed 2. cembra.
Leaves entire, their stomata ventral and dorsal 3. albicaulis.
1. PINUS KORAIENSIS
1784 P. strobus Thunberg, Fl. Jap. 275 (not Linnaeus).
1842 P. koraiensis Siebold & Zuccarini, Fl. Jap. ii. 38.
1857 P. mandschurica Ruprecht in Bull. Acad. Sci. St. Pétersb. xv. 382.
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