The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
If now the third maxilliped (Fig. 3) be examined, it will be found that,
like the swimmeret, it consists essentially of two branches springing
from a stalk of two segments. The exopodite, however, is much smaller
than the endopodite, and it ends in a flexible lash made up of many
small segments. The endopodite forms the main part of the limb, and has
five segments, so that, with the two segments of the protopodite, there
are _seven_ segments in the main axis of the limb; the second and third
segments are partly soldered together, but the line of union can be
plainly seen. Attached to the outer side of the first segment is a
membranous plate, known as the _epipodite_, on which is inserted, near
its base, a brush-like structure, which is one of the gills. In the
natural position the epipodite and its gill lie in the gill chamber,
hidden from view by the side-flap of the carapace.
[Illustration: FIG. 4--WALKING LEGS OF LOBSTER
A, Of first pair; B, of third pair]
The legs (Fig. 4) can, without difficulty, be seen to consist each of
seven segments like those of the maxillipeds, but there is no exopodite.
In the young Lobster, when just hatched from the egg, however, each of
the legs has a large exopodite like that of the third maxilliped. These
exopodites, which are used in swimming, are afterwards lost as the
animal grows; but their presence in the young is interesting as
confirming the conclusion that the legs, like the maxillipeds, are built
on the same plan as the swimmerets. The large claws, and also the first
and second pairs of walking legs, end in pincers, or _chelæ_, the
penultimate segment projecting in a thumb-like process against which the
last segment works. Each leg, except those of the last pair, has on its
first segment an exopodite with a gill like those of the maxilliped.
[Illustration: FIG. 5--APPENDAGES OF LOBSTER IN FRONT OF THIRD
MAXILLIPED
A, Eye-stalk; B, antennule; C, antenna (the flagellum is cut short); D,
mandible; E, maxillula; F, maxilla; G, first maxilliped; H, second
maxilliped. _en_, Endopodite; _ep_, epipodite; _ex_, exopodite; _gn_,
gnathobases, or jaw-plates; _p_, palp of mandible; _sc_, scaphognathite]
Following the series of appendages forwards from the third maxilliped
(Fig. 5), it is easy to trace the gradual reduction of the endopodite
and exopodite; while the two segments of the protopodite become
flattened and broadened inwards to form the jaw-plates. The mandibles
(Fig. 5, D), which are the chief organs of mastication, consist mainly
of the much enlarged basal segment of the protopodite, with a strongly
toothed inner edge, where it works against its fellow of the opposite
side; and the rest of the limb is reduced to a small sensory "palp,"
which represents the second segment of the protopodite and the
endopodite.
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