The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The antennæ (Fig. 5, C) can be shown, without difficulty, to conform to
the same plan of structure as the other appendages. The two segments of
the protopodite are short, but distinct; the endopodite forms the long
lash, or _flagellum_, composed of very numerous small segments; the
exopodite is reduced to a small movable scale or spine.
The antennules (Fig. 5, B) seem at first sight to present the
two-branched type of structure in its simplest form; but there is
considerable doubt as to whether the two lashes which each bears on a
three-segmented stalk are really equivalent to the endopodite and
exopodite.
The movable stalks which carry the eyes (Fig. 5, A) have been considered
by some to belong to the series of the appendages, and to be, in fact,
modified limbs. If this be the case, we have here the greatest
simplification which the limb undergoes in the Lobster, for each
eye-stalk consists only of two segments: the first small and
incompletely formed, the second in the form of a short cylinder, having
the eye at its end. There are, however, reasons for doubting whether the
eye-stalks are really appendages.
The hard outer covering of the Lobster not only protects and gives
support to the internal organs, but also affords points of attachment
for the muscles by means of which the animal moves. In other words, it
plays the part of a skeleton; but since, unlike the skeleton of
vertebrate animals, it is _outside_ instead of _inside_ the soft parts
of the body, it is known as an _exoskeleton_. Closer examination shows
that this outer covering is really continuous over the whole of the body
and limbs, but is thin and soft at the joints, allowing the parts to
move one upon another. It is composed of a horn-like substance known as
_chitin_, which, except at the joints, is hardened by the deposition in
it of carbonate and other salts of lime.
As this external covering does not increase in size after it has been
formed, and as it cannot stretch to any great extent, the Lobster
requires to cast its shell at intervals as it grows. In this process of
_moulting_ the integument of the back splits between the carapace and
the first abdominal somite. The body and limbs are gradually worked
loose and withdrawn through the opening, leaving the cast shell with all
its appendages almost entire. The new covering, which had been formed
underneath the old before moulting, is at first quite soft, and the
animal rapidly increases in size owing to the absorption of water. The
shell then gradually hardens by the deposition of lime salts.
[Illustration: FIG. 6--DISSECTION OF MALE LOBSTER, FROM THE SIDE. (From
British Museum Guide.)]
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