The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The internal anatomy (Fig. 6) presents many points of interest which can
only be briefly touched on here. The food-canal consists of a short
gullet leading into a capacious stomach, from which the straight
intestine runs to the vent on the under-side of the telson. The stomach
has a most remarkable and complicated structure. It consists of two
chambers, a larger in front and a smaller behind, which are lined by a
continuation of the chitinous outside covering of the body. This
chitinous lining is thickened in places to form a system of plates and
levers connected with three strong teeth set in the narrow opening
between the two chambers. By the action of muscles attached to certain
of these plates the teeth work together so as to divide up the food more
finely than had been done by the mandibles and other jaws. The whole
apparatus, in fact, serves as a kind of gizzard, and is known as the
_gastric mill_.
A small part of the intestine at the hinder end is lined, like the
stomach, by a continuation of the chitinous covering, which is turned in
at the vent. This lining and that of the stomach, with the plates and
teeth of the gastric mill, are cast and renewed when the shell is
moulted.
On each side of the food-canal in the thorax lies a large mass of soft
tissue, yellowish-green in colour. This is the _digestive gland_, or
"liver," which secretes the digestive juice, discharging it into the
food-canal by a short duct on each side just behind the stomach.
The _heart_ lies in the middle of the back, just under the hinder part
of the carapace, and gives off, in front and behind, a number of
arteries which carry the blood to the various organs of the body. From
the smaller branches of these arteries the blood passes, not, as in
vertebrate animals, into capillaries, but into the spaces lying between
the organs of the body, and it finds its way back to the heart, not in
definite veins, but by ill-defined venous channels which open into the
pericardium, or space surrounding the heart. From the pericardium the
blood enters the heart by six openings in its walls, each guarded by a
pair of valves which close when the heart contracts, and prevent the
blood from returning to the pericardium.
[Illustration: FIG. 7--GILLS OF THE LOBSTER, EXPOSED BY CUTTING AWAY THE
SIDE-FLAP OF THE CARAPACE (BRANCHIOSTEGITE)]
Public-domain text, read in full here on John Shaqi.
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