The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The subclass COPEPODA comprises animals, for the most part of
microscopic size, which are abundant in fresh water and in the sea. The
common fresh-water genus _Cyclops_ (Fig. 14) furnishes a good example of
the type of structure characteristic of the class. The body is somewhat
pear-shaped, with a narrow abdomen ending in a "caudal fork." The body
is divided into somites, and there is no overlapping carapace, although
the head and the first two thoracic somites are coalesced. There are
four pairs of two-branched, oar-like, swimming feet, and a fifth pair,
found in some other Copepoda, is represented in _Cyclops_ by minute
vestiges on the first segment of the narrow posterior part of the body.
The antennules are very large, unbranched and composed of numerous
segments; the antennæ are much smaller. In addition to the usual
mandibles, maxillulæ, and maxillæ, there is a pair of maxillipeds which
really represent the first pair of trunk limbs. There is a single red
eye in the middle of the front of the head. This eye is not formed, like
the single eye of the Cladocera, by fusion of a pair of eyes, but it
corresponds to a median eye of simple structure which is found in the
Branchiopoda, Ostracoda, and many other Crustacea, in addition to the
paired compound eyes. From the fact that this median eye is the only one
present in the earliest larval stage of Crustacea, the Nauplius (see
Chapter IV.), it is sometimes known as the "nauplius eye." The female
_Cyclops_ carries her eggs until they hatch, in two oval packets
attached to the sides of the body.
Forming a separate order (BRANCHIURA) apart from the more normal
Copepoda (order EUCOPEPODA) is the little group of the Carp-lice, one of
which, _Argulus foliaceus_, is common in England, living as a parasite
on different species of fresh-water fish, and often found swimming free
in ponds and rivers. It has a broad, flat, and very transparent body,
about three-sixteenths of an inch in length. It differs from _Cyclops_
in a great many points, of which, perhaps, the most conspicuous is the
possession of a pair of true compound eyes in addition to the median
eye. On the under-side of the head are a pair of large round suckers, by
means of which the animal fixes itself on to its prey. A study of their
development shows that these suckers are really the maxillæ, which in
the young animal are jointed limbs ending in a strong claw, but later
become changed into the suckers of the adult. A sharp spine, which can
be protruded in front of the mouth, is connected with what is believed
to be a poison-gland. The eggs are not carried in packets by the female
as in _Cyclops_, but are deposited on stones or water-weeds.
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