The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The fourth subclass, CIRRIPEDIA, comprises the Barnacles and
Acorn-shells. These are very unlike any of the other Crustacea, and, in
fact, they were long classed by naturalists with the Mollusca. It was
not until their larval development was made known that they were
recognized as Crustacea. The common Goose Barnacle (_Lepas
anatifera_--Plate III.) is found adhering to the bottoms of ships and to
floating timber. It has a fleshy stalk or peduncle which is fixed at one
end to the supporting object, and bears at the other end a shell, made
up of five separate plates, enclosing the body of the animal. The stalk
corresponds to the front part of the head, and careful examination may
discover at its end, among the hardened cement which fixes it to the
support, the remains of the antennules by which the attachment of the
young animal was first effected. The body of the animal within the
carapace or shell bears the usual mandibles, maxillulæ, and maxillæ,
close to the mouth, and six pairs of long, tendril-like feet. These feet
have each two branches, composed of numerous short segments and fringed
with long hairs. They can be protruded from the slit-like opening of the
shell, forming a sort of "casting-net" for the capture of minute
floating prey.
The Acorn-shells, of which one species (_Balanus balanoides_--Plate
III.) is abundant everywhere on our coasts, covering rocks and stones
just below high-water mark, differ from _Lepas_ and its allies in having
no peduncle. The shell is cemented directly to the rock, and is conical
in shape, like a small limpet, with a hole at the top which is closed
by four movable valves.
[Illustration: _PLATE III_
GROUP OF SPECIMENS OF THE GOOSE-BARNACLE (_Lepas anatifera_), ONE
SHOWING THE CIRRI EXTENDED AS IN LIFE. (NATURAL SIZE)
(_From Brit. Mus. Guide_)
GROUP OF A COMMON SPECIES OF ACORN-SHELL OR ROCK BARNACLE (_Balanus
balanoides_)(NATURAL SIZE)]
The Stalked Barnacles, like _Lepas_ (suborder _Pedunculata_), and the
Sessile Barnacles, or Acorn-shells, like _Balanus_ (suborder
_Operculata_), together form the order THORACICA. Of the other orders
which compose the subclass Cirripedia, the only one that need be
mentioned here is the RHIZOCEPHALA, which comprises strangely degenerate
parasites living on other Crustacea.
The Cirripedia are unlike nearly all other Crustacea in the fact that,
with few exceptions, they are hermaphrodite, having both sexes united in
each individual. In certain species of the Stalked Barnacles, however,
there are minute male individuals that are attached, like parasites, to
the large hermaphrodites. In a few species the large individuals only
possess female organs, so that the separation of the sexes is complete.
The remarkable larval metamorphoses of Cirripedes and the modifications
of structure presented by some parasitic forms will be described in
later chapters.
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