The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The orders of the Eumalacostraca, again, are grouped, as shown in the
table of classification, into four divisions. The first of these, the
SYNCARIDA, includes only one order, comprising a few small Crustacea
(see Fig. 84, p. 264) which have recently been discovered in fresh water
in Tasmania and Australia. They have no carapace, and all the thoracic
somites, or all but the first, are distinct. The antennules are
two-branched, the antennæ may have a scale-like exopodite, and the last
pair of abdominal appendages form, with the telson, a tail-fan. The eyes
are sometimes stalked, but in one species they are sessile. The thoracic
limbs, which are not clearly divided into maxillipeds and legs, carry a
double series of plate-like gills or epipodites. As will be shown later,
the living Syncarida are especially interesting on account of their
resemblance to certain very ancient fossil Crustacea.
The second division of the Eumalacostraca, the PERACARIDA, includes five
orders, the members of which differ very greatly in appearance. They all
agree, however, in certain important points of structure, of which the
most conspicuous is the possession, in the female sex, of a brood-pouch
for carrying the eggs and young. This brood-pouch is formed by a series
of overlapping plates attached to the bases of the thoracic limbs.
[Illustration: FIG. 16--_Mysis relicta_, ONE OF THE MYSIDACEA. ENLARGED.
(From Lankester's "Treatise on Zoology," after Sars.)
_cs_, Cervical groove of the carapace; _m_, brood-pouch]
The first order of the Peracarida, the MYSIDACEA, consists of small,
free-swimming, shrimp-like animals (Fig. 16). Many species are common in
the sea round the British coasts, and from their possession of a
brood-pouch, in which the young are carried, they are sometimes known as
"Opossum Shrimps." The eyes are stalked, and the carapace is well
developed, although it does not unite with all the thoracic somites. The
antennæ have a flattened, scale-like exopodite, probably of use for
keeping the animal balanced in swimming. Only one pair of the thoracic
limbs are modified to form maxillipeds, and all the legs (as in the
larval Lobster) have exopodites which form the chief swimming organs.
The uropods and telson form a "tail-fan." One of the most curious points
in the organization of some Mysidacea is the possession of a pair of
statocysts in the endopodites of the uropods. Each statocyst consists of
a small cavity containing a cake-shaped concretion known as a
"statolith," resting on a group of sensory hairs. There is reason to
believe that these organs have the same function as the statocysts of
the Lobster, although they are placed at the other end of the body. The
statolith serves the same purpose as the sand-grains found in the
Lobster's statocyst, although, unlike these, it is not introduced from
outside, but is formed in position by secretion from the walls of the
sac.
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