The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
The fifth and last subclass, that of the MALACOSTRACA, is by far the
largest and most important, and will require to be considered in more
detail than any of the others. The animals composing the various orders
into which the subclass is divided differ very greatly in structure, but
they all agree in having typically the same number of appendages as the
Lobster--namely, nineteen pairs (or twenty, if the eye-stalks be
included). They also agree in the very important character that the
trunk limbs are divided into two sets, thoracic and abdominal, the
former of eight, and the latter of six pairs.
[Illustration: FIG. 15--_Nebalia bipes._ ENLARGED. (From Lankester's
"Treatise on Zoology," after Claus.)
_a´_, Antennule; _a´´_, antenna; _ab_1-_ab_6, the abdominal limbs; _ad_,
muscle joining the two valves of the shell; _f_, tail-fork; _p_, palp of
maxillula; _r_, rostral plate; _t_, telson; 1-7, the seven somites of
the abdomen]
The first order of the Malacostraca, the NEBALIACEA, comprises a few
Crustacea of small size, which differ in some very important characters
from all the other orders. _Nebalia bipes_ (Fig. 15), which occurs on
the southern coasts of the British Isles, has a large bivalved carapace
enclosing most of the limbs. In front, a small "rostral plate" is joined
to the carapace by a movable hinge, and partly covers the stalked eyes.
The eight pairs of thoracic feet are all alike, and are flattened and
leaf-like in form, resembling those of the Branchiopoda. The first four
pairs of abdominal limbs are large two-branched swimming feet, but the
last two pairs are reduced to small vestiges. Two of the most important
points in which the Nebaliacea differ from all the other Malacostraca
are that there are seven instead of six somites in the abdomen (the last
somite has no appendages), and that the telson has connected with it a
pair of movable rods forming a "caudal fork" like that of the
Branchiopoda. On account of the leaf-like thoracic feet and the
possession of a caudal fork and other features, the Nebaliacea were
formerly classified with the Branchiopoda, but a closer examination of
their structure has shown that they are true Malacostraca. In having an
additional somite in the abdomen and in other points, however, they may
be regarded as forming a link between the Malacostraca and the lower
forms of Crustacea, and for this reason they are set apart as a series
LEPTOSTRACA, while the other orders form a series EUMALACOSTRACA.
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