Speaking broadly, evolutionists of to-day may be said to represent three
distinct lines of thought. For the sake of classification we may speak of
them as falling into three schools, which we may term the Neo-Lamarckian,
the Wallaceian, and the Neo-Darwinian, according as their views incline
towards those held by Lamarck, Wallace, or Darwin.
The Neo-Lamarckian School
As adherents of the Neo-Lamarckian school, we cite Cope, Spencer, Orr,
Eimer, Naegeli, Henslow, Cunningham, Haeckel, Korchinsky, and a number of
others. It may almost be said of these Neo-Lamarckians that each holds a
totally distinct theory of evolution. So heterogeneous are their views
that it is difficult to find a single article common to the evolutionary
belief of all. It is commonly asserted that all Neo-Lamarckians are
agreed, firstly, that acquired characters are transmissible; and,
secondly, that such transmission is an important factor in the production
of new species. This assertion is certainly true of the great bulk of
Neo-Lamarckians, but it does not appear to hold in the case of those who
believe that evolution is the result of some unknown inner force. So far
as we can see, a belief in the inheritance of acquired characters is not
necessary to the theories of orthogenesis held by Naegeli and Korchinsky.
For that reason it would possibly be more correct to place those who hold
such views in a fourth school. Since, however, a number of undoubted
Neo-Lamarckians, as, for example, Cope, believe in an inner growth-force,
it is convenient to regard Naegeli as a Neo-Lamarckian. His views need
not detain us long. Those who wish to study them in detail will find them
in his _Mechanisch-physiologische Theorie der Abstammungslehre_.
Naegeli believes that there is inherent in protoplasm a growth-force,
which makes each organism in itself a force making towards progressive
evolution. He holds that animals and plants would have become much as
they are now even if no struggle for existence had taken place. “To the
believers in this kind of . . . orthogenesis,” writes Kellog (_Darwinism
To-day_, p. 278), “organic evolution has been, and is now, ruled by
unknown inner forces inherent in organisms, and has been independent of
the influence of the outer world. The lines of evolution are immanent,
unchangeable, and ever slowly stretch toward some ideal goal.” It is easy
to enunciate such a theory, impossible to prove it, and difficult to
disprove it.
It seems to us that the fact that, so soon as organisms are removed from
the struggle for existence, they tend to degenerate, is a sufficient
reason for refusing to accept theories of the description put forth by
Naegeli. More truly Lamarckian is Eimer’s theory of orthogenesis,
according to which it is the environment which determines the direction
which variation takes; and the variations which are induced by the
environment are transmitted to the offspring.
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