Orr’s Views
Spencer and Orr preach nearly pure Lamarckism. The former, while fully
recognising the importance of natural selection, considered that
sufficient weight has not been given to the effects of use and disuse, or
to the direct action of the environment in determining or modifying
organisms.
The similarity of the views of Orr and Lamarck is best seen by comparing
their respective explanations of the long neck of the giraffe. Lamarck
thought that this was the direct result of continual stretching. The
animal continually strains its neck in the search for food, hence it
grows longer as the individual grows older, and this elongated neck has
been transmitted to the offspring. Orr writes, on page 164 of his
_Development and Heredity_: “The giraffe seems to present the most
remarkable illustration of the lengthening of the bones as the result of
the frequent repetition of such shocks. As is well known, this animal
feeds on the foliage of trees. From the earliest youth of the species,
and the earliest youth of each individual, it must have been stretching
upwards for food, and, as is the custom of such quadrupeds, it must have
constantly raised itself off its forefeet, and, as it dropped, must have
received a shock that made itself felt from the hoofs through the legs
and vertical neck to the head. In the hind legs the shock would not be
felt. It is impossible to imagine that an animal which, during the
greater part of every day of its life (both its individual and racial
life), performed motions so uniform and constant, would not be peculiarly
specialised as a result. The forces acting upon such an animal are widely
different from the forces acting upon an animal which eats the grass at
its feet like an ox, or one which must run and climb like a goat or a
deer, and the resultant modifications of growth in the several cases must
also be different. The principle of increased growth in the direction of
the shock, resulting from superabundant repair of the momentary
compression, explains how the giraffe acquired the phenomenal length of
the bones of its forelegs and neck; and the absence of the shock in the
hind-quarters shows why they remained undeveloped and absurdly
disproportionate to the rest of the body.”
Inheritance of Acquired Characters
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