8. An almost insuperable objection to the theory that species have
originated by the action of natural selection on minute variations, is
that such small differences cannot be of a life-or-death value, or, as it
is usually called, a survival value to their possessor. But if evolution
is the result of the preservation by natural selection of such slight
variations, it is absolutely necessary that each of these should possess
a survival value.
As D. Dewar has pointed out, on page 704 of vol. ii. of _The Albany
Review_, it is only when the beast of prey and its victim are evenly
matched as regards fleetness and power of endurance that small variations
in these qualities can have a survival value. But in the rough and tumble
of the struggle for existence the victim and its foe are but rarely
well-matched. Take as an example the case of a flycatcher. “This bird,”
writes D. Dewar, “will sometimes take three or four insects in the course
of one flight; all are captured with the same ease, although the length
of wing in each victim varies. So great is the superiority of the bird
that it does not notice the difference in the flying powers of its puny
quarry.” It is unnecessary to labour this point.
9. Species or varieties differing considerably in colour may exist side
by side, as the hooded and carrion crows, the white and dark breasted
forms of the Arctic skua, the pale and dark forms of the fulmar petrel,
the grey and rufous forms of the American scops owl (_Megascops asio_).
It is true that preponderance of one form or another in certain districts
points to some advantage possessed by one over the other, but, for all we
know, it may be due to heredity, and in any case the co-existence of the
two types in part of their range, or at certain seasons, shows that
selection is not at all rigorous.
The same argument applies to the co-existence of very
differently-coloured species with generally similar habits, such as that
of the jaguar and puma in South America, and the five very
differently-coloured flycatchers in the Nilgiri Hills.
Leaf-butterflies
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